The Caesar grunt (''Haemulon carbonarium''), also known as the black grunt, blacktail grunt, or redmouth grunt is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

, a grunt in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Haemulidae Haemulidae is a family of fishes in the order Perciformes known commonly as grunts. It is made up of the two subfamilies Haemulinae (grunters) and Plectorhynchinae (sweetlips), which in turn contain about 133 species in 19 genera. These fish ar ...

. It is native to the western

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

Description

The Caesar grunt has an oblong, compressed body with a deep, blunt-snouted head. The moderately sized mouth is horizontal and does not have very fleshy lips. It has a continuous

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

, which has a slight notch at its centre. The base of soft-rayed part of the dorsal fin and of the

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

are scaled. The dorsal fin contains 12 spines and 15-16 soft rays while the anal fin has tree spines and eight soft rays. The overall colour is silvery white broken by longitudinal orange-brown stripes, these are broader and more obvious above the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...

. Also, they have a black spot at the base of their

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

, which is characteristic of this species. The anal,

pectoral Pectoral may refer to: * The chest region and anything relating to it. * Pectoral cross, a cross worn on the chest * a decorative, usually jeweled version of a gorget * Pectoral (Ancient Egypt), a type of jewelry worn in ancient Egypt * Pectoralis ...

, and

pelvic fins Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...

are transparent, while the caudal fin is dark. The dorsal fin is also transparent, but it has an orange-brown margin on the spiny part, while the soft-rayed part has a dark margin. The head is marked with bronze-yellow stripes and spots. The flesh on the inside of the mouth is orange. This species attains a maximum total length of , although is more typical.

Distribution

The Caesar grunt is found in the western Atlantic Ocean. It is distributed from

Cape Canaveral , image = cape canaveral.jpg , image_size = 300 , caption = View of Cape Canaveral from space in 1991 , map = Florida#USA , map_width = 300 , type =Cape , map_caption = Location in Florida , location ...

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...

and

Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...

in the north southwards through the Bahamas, to the

Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...

including the

Florida Keys The Florida Keys are a coral cay archipelago located off the southern coast of Florida, forming the southernmost part of the continental United States. They begin at the southeastern coast of the Florida peninsula, about south of Miami, and e ...

, the

Florida Middle Grounds The Florida Middle Grounds are a reefs in the northeastern Gulf of Mexico, approximately west northwest of the West coast of Florida. Geography These reefs consist of a series of both high and low relief limestone ledges and pinnacles that exc ...

, along the mainland coast from

Tuxpan Tuxpan (or Túxpam, fully Túxpam de Rodríguez Cano) is both a municipality and city located in the Mexican state of Veracruz. The population of the city was 78,523 and of the municipality was 134,394 inhabitants, according to the INEGI census o ...

in Mexico along the northern coast of the Yucatan Peninsula to northwestern Cuba, and throughout the

Caribbean Sea The Caribbean Sea ( es, Mar Caribe; french: Mer des Caraïbes; ht, Lanmè Karayib; jam, Kiaribiyan Sii; nl, Caraïbische Zee; pap, Laman Karibe) is a sea of the Atlantic Ocean in the tropics of the Western Hemisphere. It is bounded by Mexico ...

. Claims from Brazil are likely to be misidentifications.

Habitat and biology

The Caesar grunt is found at depths from . This species is found in clear water on rocky or coral reefs and in mangroves. They are social Fish which form schools, feeding nocturnally on benthic invertebrates such as

crabs Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...

gastropods The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. Ther ...

starfish Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish ...

and

polychaetes Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are mad ...

. Caesar grunt’s are oviparous, forming distinct pairs to spawn and have pelagic larvae.

Systematics

The Caesar grunt was first formally described in 1860 by the

Cuban Cuban may refer to: * Something of, from, or related to Cuba, a country in the Caribbean * Cubans, people from Cuba, or of Cuban descent ** Cuban exile, a person who left Cuba for political reasons, or a descendant thereof * Cuban citizen, a perso ...

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

Felipe Poey (1799–1891) with the type locality given as

Havana Havana (; Spanish: ''La Habana'' ) is the capital and largest city of Cuba. The heart of the La Habana Province, Havana is the country's main port and commercial center.

. The specific name ''carbonarius'' means “coal-like”, a reference to its Cuban Spanish vernacular name of ''Ronco carbonero'', named for its dark fins.

Uses

The Caesar grunt is targeted by commercial fisheries in some parts of its range, it is also caught for the

aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...

trade. They are caught using traps, hook-and-line, and seines, but catch statistics are not collected. They are mainly sold fresh.

References

External links

* {{Taxonbar, from=Q2368941 Caesar grunt Fish of the Caribbean Fish of the Western Atlantic Caesar grunt