Haemanthinae is a small

subtribe Subtribe is a taxonomic category ranking which is below the rank of tribe and above genus. The standard suffix for a subtribe is -ina (in animals) or -inae (in plants Plants are predominantly photosynthetic eukaryotes of the kingdom Plant ...

Haemantheae Haemantheae are a tribe of subfamily Amaryllidoideae (family Amaryllidaceae). They are herbaceous monocot perennial flowering plants with a predominantly African distribution. Three subtribes are proposed and six genera including the type genus, ...

, and therefore within the African

clades A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, t ...

Amaryllidoideae Amaryllidoideae (Amaryllidaceae ''s.s.'', amaryllids) is a subfamily of monocot flowering plants in the family Amaryllidaceae, order Asparagales. The most recent APG classification, APG III, takes a broad view of the Amaryllidaceae, which then ...

. It consists of two

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

, ''

Haemanthus ''Haemanthus'' is a Southern African genus of flowering plants in the family Amaryllidaceae, subfamily Amaryllidoideae. Members of the genus are known as blood lily and paintbrush lily. There are some 22 known species, native to South Africa, B ...

'', and ''

Scadoxus ''Scadoxus'' is a genus of African and Arabian plants in the Amaryllidaceae, Amaryllis family, subfamily Amaryllidoideae. The English names blood lily or blood flower are used for some of the species. The genus has close affinities with ''Haemant ...

''.

Description

The genera of Haemanthinae share brush-like

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

s, in which the

bracts In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...

frequently form part of the pollinator attraction system. Not all ''Scadoxus'' form

bulbs In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs durin ...

, while all species of ''Haemanthus'' do.

Taxonomy

For the early taxonomic history of these two genera, see Meerow and Clayton (2004).

Pax Pax or PAX may refer to: Peace * Peace (Latin: ''pax'') ** Pax (goddess), the Roman goddess of peace ** Pax, a truce term * Pax (liturgy), a salutation in Catholic and Lutheran religious services * Pax (liturgical object), an object formerly kiss ...

(1888) treated the Amaryllidaceae as four subfamilies, with two tribes in subfamily Amarylloideae, which consisted of two tribes, Amaryllideae and Narcisseae. The former contained six subtribes, placing ''Haemanthus'' together with ''

Clivia ''Clivia'' is a genus of monocot flowering plants native to southern Africa. They are from the family Amaryllidaceae, subfamily Amaryllidoideae. Common names are Natal lily or bush lily. They are herbaceous or evergreen perennial plants, with g ...

'' in subtribe Haemanthinae. Hutchinson (1934) subsequently elevated this to

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...

Haemantheae. Later, Traub placed ''Haemanthus'' with ''Choananthus'' (subsequently submerged in ''Scadoxus'') in tribe Haemantheae in his 1963 monograph on the

Amaryllidaceae The Amaryllidaceae are a family of herbaceous, mainly perennial and bulbous (rarely rhizomatous) flowering plants in the monocot order Asparagales. The family takes its name from the genus ''Amaryllis'' and is commonly known as the amaryllis fa ...

. Later the Müller-Doblies' created a narrower concept of Haemantheae as a tribe with Haemanthinae as one of two subtribes and two genera, ''Haemanthus'' and ''Scadoxus''.

Molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

research has confirmed this placement, with

Meerow Alan W. Meerow is an American botanist, born in New York City in 1952. He specializes in the taxonomy of the family Amaryllidaceae and the horticulture of palms and tropical ornamental plants. He also works on the population genetics and mole ...

and Clayton (2004) situating Haemanthinae as one of three subtribes of Haemantheae.

Phylogeny

The Haemanthinae are placed within the Haemantheae as follows:

Subdivision

* ''

'' (21 species) * ''

'' (9–12 species)

Distribution and habitat

''Scadoxus'' is a forest

understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

genus, most commonly found in the

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

areas of Africa. Haemanthus on the other hand is confined to southern Africa, within the summer and winter rainfall regions of the Cape.

Notes

References

Bibliography

* Volume 1:

Monocotyledon Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...

ae 1926, Volume 2:

Dicotyledon The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...

ae 1934. * * * * * * * * , in * *

External links

{{Taxonbar, from=Q23012367 Amaryllidoideae Garden plants of Southern Africa Flora of Southern Africa Flora of Africa Plant subtribes