The Guayanan Highlands moist forests (NT0124) is an
ecoregion
An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of ...
in the south of Venezuela and the north of Brazil and in Guyana, Suriname, French Guiana. It is in the
Amazon biome
The Amazon biome ( pt, Bioma Amazônia) contains the Amazon rainforest, an area of tropical rainforest, and other ecoregions that cover most of the Amazon basin and some adjacent areas to the north and east. The biome contains blackwater and white ...
.
It encompasses an upland region with diverse fauna and flora, which contains dramatic
tepui
A tepui , or tepuy (), is a table-top mountain or mesa found in South America, especially in Venezuela and western Guyana. The word tepui means "house of the gods" in the native tongue of the Pemon, the indigenous people who inhabit the Gra ...
s, or sandstone table mountains.
The region has been inaccessible in the past and is generally fairly intact, apart from the north and northeast where large scale agriculture, ranching and mining operations are steadily encroaching on the ecosystem. New roads are opening the interior to logging, and planned dams will have a drastic impact on the
riparian zone
A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
s.
Location
The ecoregion includes parts of southern Venezuela, western and southern Guyana and northern Brazil, with scattered portions in Suriname and French Guiana.
It extends into eastern Colombia.
It has a total area of .
The ecoregion lies on the
Guiana Shield
The Guiana Shield (french: Plateau des Guyanes, Bouclier guyanais; nl, Hoogland van Guyana, Guianaschild; pt, Planalto das Guianas, Escudo das Guianas; es, Escudo guayanés) is one of the three cratons of the South American Plate. It is a ...
, an ancient upland area between the Amazon and Orinoco basins.
It is surrounded by lowland grassy savannas and lowland forest.
All areas of the ecoregion contains enclaves of the Pantepuis ecoregion on the tops of table mountains.
Most areas of the ecoregion in the east are surrounded by the Guianan moist forests ecoregion, and most areas in the west are surrounded by the
Guianan piedmont and lowland moist forests
The Guianan piedmont and lowland moist forests (NT0182) is an ecoregion in the south of Venezuela and the north of Brazil. It is in the Amazon biome.
The ecoregion is relatively intact, largely protected by conservation units or indigenous territor ...
ecoregion.
Sections of the ecoregion in the east border the Uatuma-Trombetas moist forests to the south.
The central part of the ecoregion surrounds the northern part of the largest section of the Guianan savanna ecoregion
The southwestern part of the ecoregion rises above the Negro-Branco moist forests ecoregion.
The Guayanan Highlands moist forests and the Tepuis together make up the Guayanan Highlands Forests
Global 200
The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature ( WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or w ...
ecoregion.
Physical
The terrain is rugged, with elevations in the ecoregion from above sea level.
Taller table mountains in the region rise to elevations of and host the
Tepui
A tepui , or tepuy (), is a table-top mountain or mesa found in South America, especially in Venezuela and western Guyana. The word tepui means "house of the gods" in the native tongue of the Pemon, the indigenous people who inhabit the Gra ...
ecoregion.
The Guayanan Highlands ecoregion is an "island" of higher land surrounded by lower grasslands and forests collectively known as the
Guiana Shield
The Guiana Shield (french: Plateau des Guyanes, Bouclier guyanais; nl, Hoogland van Guyana, Guianaschild; pt, Planalto das Guianas, Escudo das Guianas; es, Escudo guayanés) is one of the three cratons of the South American Plate. It is a ...
.
Most of the land drains into the
Orinoco
The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wor ...
Uraricoera
The Uraricoera River (Uraricuera) is a river of Roraima state in northern Brazil. The confluence of the Uraricoera and Takutu Rivers forms the Branco River
The Branco River ( pt, Rio Branco; Engl: ''White River'') is the principal affluent of t ...
Amazon
Amazon most often refers to:
* Amazons, a tribe of female warriors in Greek mythology
* Amazon rainforest, a rainforest covering most of the Amazon basin
* Amazon River, in South America
* Amazon (company), an American multinational technolog ...
.
The upland terraces and mountains consist mainly of quartzitic or sandstone rocks, with granitic rock in some areas.
The lowland plains are recent, formerly flooded by lakes or by the sea.
Most of the landscape is forest-covered rolling
peneplain
390px, Sketch of a hypothetical peneplain formation after an orogeny.
In geomorphology and geology, a peneplain is a low-relief plain formed by protracted erosion. This is the definition in the broadest of terms, albeit with frequency the usage ...
s and
floodplain
A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...
s.
The undulating peneplains between the Caura and Paragua rivers hold low hills up to high.
There are scattered upland areas of undulating plains, rounded hills, low mountains and Tepuis up to high, and the lower slopes of the higher tepuis.
Soils are mostly sandy and low in nutrients.
Ecology
The Guianan Highlands moist forests ecoregion is in the
Neotropical realm
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone.
Definition
In bioge ...
and the
tropical and subtropical moist broadleaf forests
Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature.
Description
TSMF is generally found in large, discon ...
biome.
Climate
The
Köppen climate classification
The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, nota ...
is "Am": equatorial, monsoonal.
Temperatures vary little throughout the year.
Yearly average temperatures range from a minimum of to a maximum of , with a mean of .
Annual rainfall varies considerably, but in some areas averages .
Monthly rainfall is lowest in January at and highest in July at .
Flora
There are various habitats including large areas of tall primary rainforest, open savannas and gallery forests.
The peneplains hold evergreen forests with dense canopies of with emergent trees in the
Calophyllum
''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands.
History
Members of the genus ''Calophyllu ...
,
Anacardium
''Anacardium'', the cashews, are a genus of flowering plants in the family Anacardiaceae, native to tropical regions of the Americas. The best known species is '' Anacardium occidentale,'' which is commercially cultivated for its cashew nuts and ...
Inga
''Inga'' is a genus of small tropical, tough-leaved, nitrogen-fixing treesElkan, Daniel. "Slash-and-burn farming has become a major threat to the world's rainforest" ''The Guardian'' 21 April 2004 and shrubs, subfamily Mimosoideae. ''Inga''s ...
,
Parkia
''Parkia'' is a genus of flowering plants in the family Fabaceae. It belongs to the mimosoid clade of the subfamily Caesalpinioideae. Several species are known as African locust bean.
In 1995, about 31 species were known.Melissa Luckow and H ...
,
Copaifera
''Copaifera'' is a genus of tropical plants in the legume family Fabaceae.
The scientific name means " copal-bearer" (or more accurately, ''copaiba''-bearer), since economically important resins and essential oils can be acquired from them. ...
,
Erythrina
''Erythrina'' is a genus of plants in the pea family, Fabaceae. It contains about 130 species, which are distributed in tropical and subtropical regions worldwide. They are trees, with the larger species growing up to in height. The generic n ...
and Dipteryx genera.
Common trees on the plains include ''
Micropholis melinoniana
''Micropholis'' is group of trees in the family Sapotaceae, described as a genus in 1891. (2001): World Checklist of Sapotaceae &ndash''Micropholis'' The Board of Trustees of the Royal Botanic Gardens, Kew. Retrieved 2008-DEC-24.
These trees ar ...
'',
Dacryodes
''Dacryodes'' is a genus of about 60 species of trees in the family Burseraceae. The generic name is from the Greek ' meaning "tear(drop)", referring to how resin droplets form on the bark surface.
Description
''Dacryodes'' species grow as shrub ...
species, ''
Euterpe precatoria
''Euterpe precatoria'' is a tall, slender-stemmed, pinnate-leaved palm native to Central and South America and Trinidad and Tobago. ''E. precatoria'' is used commercially to produce fruits, although '' Euterpe oleracea'' is more commonly cultiv ...
Alexa
Alexa may refer to: Technology
*Amazon Alexa, a virtual assistant developed by Amazon
* Alexa Internet, a defunct website ranking and traffic analysis service
* Arri Alexa, a digital motion picture camera
People
* Alexa (name), a given name a ...
species, ''
Euterpe precatoria
''Euterpe precatoria'' is a tall, slender-stemmed, pinnate-leaved palm native to Central and South America and Trinidad and Tobago. ''E. precatoria'' is used commercially to produce fruits, although '' Euterpe oleracea'' is more commonly cultiv ...
'' and ''
Micrandra minor
''Micrandra'' is a plant genus of the family Euphorbiaceae first described in 1854. It is native to South America.
Species
Species currently included in ''Micrandra'':
# '' Micrandra australis'' (R.E.Schult.) R.E.Schult. - Amazonas in ...
''.
The seasonally flooded forests along the rivers have similar formations of flora to the Amazon region.
Common tree species on the upper Orinoco include '' Combretum frangulifolium'', '' Gustavia augusta'',
Pterocarpus
''Pterocarpus'' is a pantropical genus of trees in the family Fabaceae. It belongs to the subfamily Faboideae, and was recently assigned to the informal monophyletic ''Pterocarpus'' clade within the Dalbergieae. Most species of ''Pterocarp ...
Inga
''Inga'' is a genus of small tropical, tough-leaved, nitrogen-fixing treesElkan, Daniel. "Slash-and-burn farming has become a major threat to the world's rainforest" ''The Guardian'' 21 April 2004 and shrubs, subfamily Mimosoideae. ''Inga''s ...
Panopsis rubescens
''Panopsis'' is a genus of trees in the family Proteaceae. The species, which occur in Central and South America, include: Newer species that can be considered part of the ''Panopsis'' genus has been discovered recently, called ''Panopsis magnifr ...
''.
Fauna
209 species of mammals have been recorded, including
jaguar
The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus ''Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
(''Panthera onca''),
cougar
The cougar (''Puma concolor'') is a large cat native to the Americas. Its range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mammal in the Western Hemisphere. ...
(''Puma concolor''),
South American tapir
The South American tapir (''Tapirus terrestris''), also commonly called the Brazilian tapir (from the Tupi ''tapi'ira''), the Amazonian tapir, the maned tapir, the lowland tapir, the ''anta'' ( Portuguese), and ''la sachavaca'' (literally "bushc ...
(''Tapirus terrestris''),
white-lipped peccary
The white-lipped peccary (''Tayassu pecari'') is a species of peccary found in Central and South America and the only member of the genus ''Tayassu''. Multiple subspecies have been identified. White-lipped peccaries are similar in appearance ...
(''Tayassu pecari''),
collared peccary
The collared peccary (''Dicotyles tajacu'') is a species of artiodactyl (even-toed) mammal in the family Tayassuidae found in North, Central, and South America. It is the only member of the genus ''Dicotyles''. They are commonly referred to as ...
(''Pecari tajacu''),
brocket deer
Brockets or brocket deer are the species of deer in the genus ''Mazama''. They are medium to small in size, and are found in the Yucatán Peninsula, Central and South America, and the island of Trinidad. Most species are primarily found in fore ...
(Mazama genus),
white-eared opossum
The white-eared opossum (''Didelphis albiventris''), known as the timbu in Brazil and comadreja overa in Argentina, is an opossum species found in Argentina, Bolivia, Brazil, Paraguay, and Uruguay. It is a terrestrial and, sometimes, arboreal ani ...
(''Didelphis albiventris''),
big lutrine opossum
The big lutrine opossum (''Lutreolina crassicaudata''), also known as the little water opossum, comadreja colorada, thick-tailed opossum, or coligrueso is an opossum species from South America in the genus ''Lutreolina''.
Description
The big ...
(''Lutreolina crassicaudata''),
Robinson's mouse opossum
Robinson's mouse opossum (''Marmosa robinsoni'') is a species of opossum in the family Didelphidae. It is found in Belize, Colombia, Ecuador, Grenada, Honduras, Panama, Peru, Trinidad and Tobago, and Venezuela.
Robinson's mouse opossums move a ...
Fernandez's sword-nosed bat
Fernandez's sword-nosed bat (''Lonchorhina fernandezi'') is a species of bat in the family Phyllostomidae. It is the smallest species of the '' Lonchorhina'' genus. It is endemic to Venezuela. In 2013, Bat Conservation International listed this ...
(''Lonchorhina fernandezi''),
highland yellow-shouldered bat
The highland yellow-shouldered bat (''Sturnira ludovici'') is a species of bat in the family Phyllostomidae. It is found in Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, and Venezuela
...
(''Sturnira ludovici''),
eastern lowland olingo
The eastern lowland olingo (''Bassaricyon alleni'') is a species of olingo from South America, where it is known from the lowlands east of the Andes in Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru and Venezuela.fiery squirrel (''Sciurus flammifer''),
Guyanan spiny rat
The Guyanan spiny-rat (''Proechimys hoplomyoides'') is a spiny rat species found in Brazil, Guyana and Venezuela. The species was first described by George Henry Hamilton Tate
George Henry Hamilton Tate (April 30, 1894 – December 24, 19 ...
(''Proechimys hoplomyoides'') and
Orinoco agouti
The Orinoco agouti (''Dasyprocta guamara'') is a species of rodent in the family Dasyproctidae. It is endemic to Delta Amacuro in Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivari ...
(''Dasyprocta guamara'').
Endangered mammals include
black bearded saki
The black bearded saki (''Chiropotes satanas'') is a species of New World monkey, native to the Amazon rainforest of South America, specifically to an area of north-eastern Brazil. It is one of five species of bearded saki. Bearded sakis are me ...
(''Chiropotes satanas''),
Fernandez's sword-nosed bat
Fernandez's sword-nosed bat (''Lonchorhina fernandezi'') is a species of bat in the family Phyllostomidae. It is the smallest species of the '' Lonchorhina'' genus. It is endemic to Venezuela. In 2013, Bat Conservation International listed this ...
(''Lonchorhina fernandezi'') and giant otter (''Pteronura brasiliensis'').
Snakes include fer-de-lance (''
Bothrops asper
Terciopelo (''Bothrops asper'') is a species of pit vipers occurring at low to moderate elevations in northeast Mexico and Central America, and into South America where it is known from elevations up to 2600 meters in the Colombian and Ecuado ...
''), palm pit-vipers (
Bothriechis
''Bothriechis'' is a genus of venomous pit vipers, commonly called palm vipersMehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . or palm-pitvipersCampbell JA, Lamar WW. 2004. ''The Venomous Rept ...
genus), coral snakes (
Micrurus
''Micrurus'' is a genus of venomous coral snakes of the family Elapidae.
Geographic range
Species in the genus ''Micrurus'' are endemic to the Americas.
Species
The following 83 species are recognized as being valid.
*'' Micrurus albicinctus'' ...
genus), boa constrictors (''
Boa constrictor
The boa constrictor (scientific name also ''Boa constrictor''), also called the red-tailed boa, is a species of large, non-venomous, heavy-bodied snake that is frequently kept and bred in captivity. The boa constrictor is a member of the family B ...
'') and bushmasters (''
Lachesis muta
''Lachesis muta'', also known as the Southern American bushmaster or Atlantic bushmaster, is a venomous pit viper species found in South America, as well as the island of Trinidad in the Caribbean. Two subspecies are currently recognized, includi ...
''). There are many green iguanas (''Iguana iguana'') and tegus lizards ( Tupinambis genus).
Endangered amphibians include the demonic poison frog (''
Minyobates steyermarki
''Minyobates steyermarki'' (formerly ''Dendrobates steyermarki'') is a species of frog in the family Dendrobatidae endemic to Cerro Yapacana in southern Venezuela. It is also known by the common names of demonic poison frog, demonic poison-arro ...
'').
631 species of birds have been recorded, including
white-cheeked pintail
The white-cheeked pintail (''Anas bahamensis''), also known as the Bahama pintail or summer duck, is a species of dabbling duck. It was first described by Carl Linnaeus in his landmark 1758 10th edition of ''Systema Naturae'' under its current s ...
(''Anas bahamensis''),
aplomado falcon
The aplomado falcon (''Falco femoralis'') is a medium-sized falcon of the Americas. The species' largest contiguous range is in South America, but not in the deep interior Amazon Basin. It was long known as ''Falco fusco-coerulescens'' or ''Fal ...
(''Falco femoralis''),
brown-throated parakeet
The brown-throated parakeet (''Eupsittula pertinax''), also known as the Prikichi, St. Thomas conure or the brown-throated conure, in aviculture, is a species of parrot in the family Psittacidae.
Taxonomy
The brown-throated parakeet was formal ...
burrowing owl
The burrowing owl (''Athene cunicularia''), also called the shoco, is a small, long-legged owl found throughout open landscapes of North and South America. Burrowing owls can be found in grasslands, rangelands, agricultural areas, deserts, or an ...
(''Athene cunicularia''), emeralds and hummingbirds in the
Amazilia
''Amazilia'' is a hummingbird genus in the subfamily Trochilinae. It is found in tropical Central and South America.
Taxonomy
The genus ''Amazilia'' was introduced in 1843 by the French naturalist René Lesson. Lesson had used ''amazilia'' ...
two-banded warbler
The two-banded warbler (''Myiothlypis bivittata'') is a species of bird in the family Parulidae. It is found in Argentina, Bolivia, Brazil, Guyana, Peru, and Venezuela. Its natural habitats are subtropical or tropical moist montane forest
...
World Wildlife Fund
The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the W ...
classes the ecoregion as "Relatively Stable/Intact".
The interior forests have been inaccessible in the past and are generally intact, although roads are being built that open them up to logging.
Logging is illegal in the state of Amazonas, Venezuela, but continues elsewhere.
Legal and illegal mining for gold, diamonds, bauxite, and iron ore are a serious threat.
Near urban centers in the north and northeast of the ecoregion the habitat is steadily being destroyed by large-scale agriculture, cattle ranching and mining.
Planned hydroelectric dams on the Caroni and Paragua rivers would huge areas with devastating effect on the riparian habitats.
Part of
UNESCO
The United Nations Educational, Scientific and Cultural Organization is a List of specialized agencies of the United Nations, specialized agency of the United Nations (UN) aimed at promoting world peace and security through international coope ...
's
Alto Orinoco-Casiquiare Biosphere Reserve
The Alto Orinoco-Casiquiare Biosphere Reserve is a UNESCO biosphere reserve in the Venezuelan Amazon biome.
Location
The Alto Orinoco-Casiquiare Biosphere Reserve was designated in 1993.
It is located between 00°30' to 04°40'N and 62°45' to 6 ...
Canaima National Park
Canaima National Park ( es, Parque Nacional Canaima) is a park in south-eastern Venezuela that roughly occupies the same area as the Gran Sabana region. It is located in Bolívar State, reaching the borders with Brazil and Guyana.
History
Ca ...
, and
Kanuku Mountains
The Kanuku Mountains are a group of mountains in Guyana, located in the Upper Takutu-Upper Essequibo region. The name means 'forest' in the Wapishana language, a reference to the rich diversity of wildlife found there. The Eastern Kanuku Mountain ...
.
The Canaima National Park protects an area of .
There are other national parks, but they are not well funded.
The
Pico da Neblina National Park
Pico da Neblina National Park ( pt, Parque Nacional do Pico da Neblina) is a national park in the state of Amazonas in the north of Brazil, bordering on Venezuela. It overlaps with several indigenous territories, which creates tensions over land ...
in Brazil and the Parima Tapirapecó and Serranía de la Neblina national parks in Venezuela cover in total.
There are many smaller conservation units in the region, but they mainly protect the upper levels of the tepuis.