Greater Banded Hornet
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''Vespa tropica'', the greater banded hornet, is a tropical species of
hornet Hornets (insects in the genus ''Vespa'') are the largest of the eusocial wasps, and are similar in appearance to their close relatives yellowjackets. Some species can reach up to in length. They are distinguished from other vespine wasps by th ...
found in Southern Asia, New Guinea and west Africa, and which has recently been discovered to be an invasive species on the Pacific island of Guam. It is a predator of
paper wasp Paper wasps are vespid wasps that gather fibers from dead wood and plant stems, which they mix with saliva, and use to construct nests made of gray or brown papery material. Some types of paper wasps are also sometimes called umbrella wasps, d ...
s and possesses a potent sting, which can cause extreme pain and swelling.


Description

The workers of ''Vespa tropica'' are about in length, while queens grow to . The head is dark brown/red; the abdomen is black with a distinct yellow stripe which covers most of the second abdominal segment. However, there is some variation across its range and in
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, bor ...
and southeastern Asia, specimens are often completely black and larger in size, while in other regions such as
Hong Kong Hong Kong ( (US) or (UK); , ), officially the Hong Kong Special Administrative Region of the People's Republic of China (abbr. Hong Kong SAR or HKSAR), is a city and special administrative region of China on the eastern Pearl River Delta i ...
, the head and flanks of the thorax are normally reddish. A third colour form is found in the
Andamans The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between the ...
and
Nicobars The Nicobar Islands are an archipelagic island chain in the eastern Indian Ocean. They are located in Southeast Asia, northwest of Aceh on Sumatra, and separated from Thailand to the east by the Andaman Sea. Located southeast of the Indian s ...
, which has a reddish brown head and thorax and all the dorsal plates on each segment of the gastrum are orange except the first.


Distribution

''Vespa tropica'' is distributed throughout southern Asia from
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
to the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
, south to
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
, New Britain and the
Torres Strait Islands The Torres Strait Islands are a group of at least 274 small islands in the Torres Strait, a waterway separating far northern continental Australia's Cape York Peninsula and the island of New Guinea. They span an area of , but their total land ...
but it has not been recorded in mainland Australia.


Invasive

In August 2016 ''Vespa tropica'' was found at two sites in
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
, and has since been deemed a permanent resident of the island's ecosystem, until a solution is found to exterminate them.


Biology

''Vespa tropica'' are raiders of the nests of other wasp species and take captured
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e back to their own nest to feed to their larvae. They have been observed to catch
honeybee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosm ...
s quite frequently, and even
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threa ...
have been noted as prey. A worker of ''V. tropica'' has been recorded as capturing and killing an individual of '' Vespa bicolor'' and also raiding nests of other polister wasps such as ''
Ropalidia marginata ''Ropalidia marginata'' is an Old World species of paper wasp. It is primitively eusocial, not showing the same bias in brood care seen in other social insects with greater asymmetry in relatedness. The species employs a variety of colony foundi ...
''. ''Vespa tropica'' is associated with forests in the lowlands, up to altitude. The nests are usually within above the ground in tree hollows or in subterranean cavities normally down to a depth of including a record of one found inside a dead log partly buried in the soil. In the vicinity of human habitations it will nest under roofs, in attics and sheds. The nest is normally rhomboid or bowl-shaped and has an open bottom, whereas most aerial hornets' nest are completely sealed. The outer shell of the nest consists of distinct, broad layers and the nest is quite brittle. Occasionally nests are built within the crown of trees. Nest temperature is maintained at until about the middle of October, when it trends towards ambient as the nest begins to die. Predation by ''V. tropica'' is one of the foremost causes of colony failure in '' Polistes chinensis'' in Japan. ''V. tropica'' may play a significant role in controlling Asian polistine populations. It can be a secretive species, preferring to fly close to cover and often high among the crowns of trees. ''V. tropica'', '' V. ducalis'', and '' V. philippinensis'' together form the sub-genus ''V. tropica'' group.


References

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External links


Vespa tropica
* {{Taxonbar, from=Q1940411 Vespidae Hymenoptera of Asia Hymenoptera of Oceania Insects of Southeast Asia Insects of Afghanistan Insects of Borneo Insects of Brunei Insects of India Insects of Indonesia Insects of Malaysia Insects of Pakistan Insects of Papua New Guinea Insects of Thailand Insects described in 1758 Taxa named by Carl Linnaeus