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A goanna is any one of several
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of lizards of the genus '' Varanus'' found in Australia and Southeast Asia. Around 70 species of ''Varanus'' are known, 25 of which are found in Australia. This varied group of
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s ranges greatly in size and fills several
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...
s. The goanna features prominently in
Aboriginal mythology Australian Aboriginal religion and mythology is the sacred spirituality represented in the stories performed by Aboriginal Australians within each of the language groups across Australia in their ceremonies. Aboriginal spirituality includes ...
and
Australian folklore Australian folklore refers to the folklore and urban legends that have evolved in Australia from Aboriginal Australian myths to colonial and contemporary folklore including people, places and events, that have played part in shaping the culture, ...
. Being
predatory Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
lizards, goannas are often quite large, or at least bulky, with sharp teeth and claws. The largest is the
perentie The perentie (''Varanus giganteus'') is the largest monitor lizard or goanna native to Australia. It is one of the largest living lizards on earth, after the Komodo dragon, Asian water monitor, crocodile monitor, and intersecting by size with ...
(''V. giganteus''), which can grow over in length. Not all goannas are so large; pygmy goannas may be smaller than the arm of an adult human. The smallest of these, the short-tailed monitor (''V. brevicauda''), reaches only in length. They survive on smaller prey, such as insects and mice. Goannas combine predatory and scavenging behaviours. They prey on any animal they can catch that is small enough to eat whole. They have been blamed by farmers for the death of
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
, though most likely erroneously, as goannas are also eaters of
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
and are attracted to rotting meat. Most goannas are dark-coloured, with greys, browns, blacks, and greens featuring prominently; however, white is also common. Many desert-dwelling species also feature yellow-red tones.
Camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
ranges from bands and stripes to splotches, speckles, and circles, and can change as the creature matures, with juveniles sometimes being brighter than adults. Like most lizards, goannas lay
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
. Most lay eggs in a nest or burrow, but some species lay their eggs inside
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
mounds. This offers protection and incubation; additionally, the termites may provide a meal for the young as they hatch. Unlike some other species of lizards, goannas do not have the ability to regrow limbs or tails.


Etymology

The name goanna derived from
iguana ''Iguana'' (, ) is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described in 1768 by Austrian naturalist Josephus Nicolaus Laurenti in his bo ...
, since early European bush settlers in Australia likened goannas to the South American lizards. Over time, the initial vowel sound was dropped. A similar explanation is used to link possums to the American
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 93 species in 18 genera. Opossums originated in South America and entered North ...
. Another possibility is that the name might have derived from the South African term for a monitor lizard, ''
leguaan The Nile monitor (''Varanus niloticus'') is a large member of the monitor family (Varanidae) found throughout most of Sub-Saharan Africa and along the Nile, with invasive populations in North America. The population in West African forests and ...
'', as the
Cape of Good Hope The Cape of Good Hope ( af, Kaap die Goeie Hoop ) ;''Kaap'' in isolation: pt, Cabo da Boa Esperança is a rocky headland on the Atlantic coast of the Cape Peninsula in South Africa. A common misconception is that the Cape of Good Hope is t ...
was a popular refresher stop for immigrant ships to Australia from Britain. Ironically, the term ''leguaan'' is also from Spanish 'le iguana'.


Species

For a list of all monitor lizards of the genus, see Complete list of genus ''Varanus''. The following are found in Australia. For the most part, in common names, "goanna" and "monitor" are interchangeable. *
Perentie The perentie (''Varanus giganteus'') is the largest monitor lizard or goanna native to Australia. It is one of the largest living lizards on earth, after the Komodo dragon, Asian water monitor, crocodile monitor, and intersecting by size with ...
– ''V. giganteus'' *
Lace monitor The lace monitor or tree goanna (''Varanus varius'') is a member of the monitor lizard family native to eastern Australia. A large lizard, it can reach in total length and in weight. The lace monitor is considered to be a least-concern species ...
– ''V. varius'' *
Sand goanna The sand goanna (''Varanus gouldii'') is a species of large Australian monitor lizard, also known as Gould's monitor, sand monitor, or racehorse goanna. Taxonomy John Edward Gray described the species in 1838 as ''Hydrosaurus gouldii'', noting ...
&nd arttu on homoash; ''V. gouldii'' (also Gould's goanna or ground goanna) *
Mertens' water monitor Mertens' water monitor (''Varanus mertensi''), also called commonly Mertens's water monitor, and often misspelled Mertin's water monitor, is a species of lizard in the family Varanidae. The species is endemic to northern Australia, and is a wide ...
– ''V. mertensi'' *
Spiny-tailed monitor The spiny-tailed monitor (''Varanus acanthurus''), also known as the Australian spiny-tailed monitor, the ridge-tailed monitor the Ackie's dwarf monitor, and colloquially simply ackie monitor, is an Australian species of lizard belonging to the g ...
– ''V. acanthurus'' (also ridge-tailed monitor) *
Mangrove monitor The mangrove monitor, mangrove goanna, or Western Pacific monitor lizard (''Varanus indicus'') is a member of the monitor lizard family with a large distribution from northern Australia and New Guinea to the Moluccas and Solomon Islands. It grows ...
– ''V. semiremex'' *
Black-headed monitor The black-headed monitor or black-tailed monitor (''Varanus tristis'') is a relatively small species of monitor lizards native to Australia. It is occasionally also called the mournful monitor, freckled monitor (''Varanus tristis orientalis'') o ...
– ''V. tristis'' * Short-tailed monitor – ''V. brevicauda'' *
Argus monitor The yellow-spotted monitor (''Varanus panoptes''), also known as the Argus monitor, is a monitor lizard found in northern and western regions of Australia and southern New Guinea. Taxonomy The first description of the species was provided in a ...
– ''V. panoptes'' (also yellow-spotted monitor) * Rosenberg's monitor – ''V. rosenbergi'' *
Spencer's goanna Spencer's goanna (''Varanus spenceri''), also known commonly as Spencer's monitor, is a species of monitor lizard in the family Varanidae. The species is endemic to Australia. Etymology The specific name, ''spenceri'', is in honor of Englis ...
– ''V. spenceri'' *
Storr's monitor There are two species of lizard named Storr's monitor: * ''Varanus storri Storr's monitor (''Varanus storri'') is a species of monitor lizard in the family Varanidae. The species is endemic to Australia. Etymology The specific name, ''storri ...
– ''V. storri'' *
Dampier Peninsula monitor The Dampier Peninsula monitor or Dampier Peninsula goanna (''Varanus sparnus''), described in 2014, is the smallest known species of monitor lizard, growing up to 16.3 grams with a length of almost 23 cm and a SVL (snout to vent length) of ...
– ''V. sparnus'' * Mitchell's water monitor – ''V. mitchelli'' * Kings' monitor – ''V. kingorum'' * Southern Pilbara rock goanna – ''V. hamersleyensis'' *
Black-palmed rock monitor The black-palmed rock monitor (''Varanus glebopalma'') is a member of the Varanidae family found in Australia. Also known as the twilight monitor or the long-tailed rock monitor, it is a member of the subgenus ''Odatria'', and is found in the n ...
– ''V. glebopalma'' *
Kimberley rock monitor The Kimberley rock monitor (''Varanus glauerti'') is a medium-sized species of monitor lizard
– ''V. glauerti'' *
Pygmy mulga monitor The pygmy mulga monitor (''Varanus gilleni''), also known as Gillen's monitor or just mulga monitor is a species of lizard in the family Varanidae. Taxonomy A description of the species was presented to the Royal Society of Victoria in 1895 by ...
– ''V. gilleni'' *
Rusty desert monitor The rusty desert monitor (''Varanus eremius'') is a species of small monitor lizards native to Australia. It is also known as the pygmy desert monitor. The monitor lizard belongs to the subgenus ''Odatria'' along with the Varanus gilleni, pygmy m ...
– ''V. eremius'' * Stripe-tailed goanna – ''V. caudolineatus'' * Pilbara monitor – ''V. bushi'' * Black-spotted ridge-tailed monitor – ''V. baritji'' *
Emerald tree monitor The emerald tree monitor (''Varanus prasinus'') or green tree monitor, is a small to medium-sized arboreal monitor lizard. It is known for its unusual coloration, which consists of shades from green to turquoise, topped with dark, transverse d ...
– ''V. prasinus'' *
Canopy goanna ''Varanus keithhornei'', commonly known as the canopy goanna, Keith Horne's monitor, blue-nosed tree monitor, or Nesbit River monitor, is a species of monitor lizards native to northeast Australia. It is a member of the '' Varanus prasinus'' ...
– ''V. keithhornei''


Habitats

Goannas are found throughout most of Australia, except for
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
, and manage to persist in a variety of environments. Most species are known to climb trees or outcrops; several primarily
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
species are known. The
lace monitor The lace monitor or tree goanna (''Varanus varius'') is a member of the monitor lizard family native to eastern Australia. A large lizard, it can reach in total length and in weight. The lace monitor is considered to be a least-concern species ...
(''V. varius'') is probably the best-known among these, but is not the most common. The lace monitor is the second-largest of all goannas, reaching lengths up to . Other more common tree goannas, such as the Timor tree monitor (''V. timorensis'') and mournful tree monitor (''V. tristis'',) do not grow to quite such lengths, typically a maximum of 61 cm, nose-to-tail. Other goannas are adapted to swampy coastal environments, such as the
mangrove goanna The mangrove monitor, mangrove goanna, or Western Pacific monitor lizard (''Varanus indicus'') is a member of the monitor lizard family with a large distribution from northern Australia and New Guinea to the Moluccas and Solomon Islands. It gro ...
(''V. semiremex''). Further still,
Mertens' water monitor Mertens' water monitor (''Varanus mertensi''), also called commonly Mertens's water monitor, and often misspelled Mertin's water monitor, is a species of lizard in the family Varanidae. The species is endemic to northern Australia, and is a wide ...
(water goanna – ''V. mertensi''), found in lagoons and rivers across northern Australia, is streamlined for swimming, using its tail as a paddle. Most other goannas are good swimmers, but tend not to voluntarily venture into water.


Diet

The diets of goannas vary greatly depending on the species and the habitat. Prey can include all manner of small animals: insects, smaller lizards, snakes, mammals, birds, and eggs. Meals are often eaten whole, thus the size of their meals may depend on the size of the animals. Many of the small species feed mostly on insects, with some being small lizard experts. Many of the medium to large species feed on whatever prey they can catch. This includes eggs, fish (''V. mertensi''), birds, snails, smaller lizards, snakes, marsupials, and other small mammals, such as rodents. The giant perentie has been observed killing a young
kangaroo Kangaroos are four marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...
, and then biting out chunks of flesh like a dog. All species are carrion eaters, so feed on the carcasses of dead animals, including livestock and other large creatures. The smell of rotting meat also attracts these lizards.


Goannas and humans


Confrontations

Like most native fauna, goannas are rather wary of human intrusions into their habitat, and most likely run away (into the scrub, up a tree, or into the water, depending on the species). A goanna is a rather swift mover, and when pressed, sprints short distances on its hind legs. Goannas also rear up when threatened, either chased or cornered, and also inflate flaps of skin around their throats and emit harsh hissing noises. Some goannas lose their initial fear of humans, especially when food is involved (or has been previously involved). The wildlife authority recommends not feeding animals while in their territory. An attack can cause serious injury in exceptional cases, but most authorities doubt that a goanna will direct an intentional attack at a human unless the human has attempted to attack it (or grasp at it) first.
Indigenous Australians Indigenous Australians or Australian First Nations are people with familial heritage from, and membership in, the ethnic groups that lived in Australia before British colonisation. They consist of two distinct groups: the Aboriginal peoples ...
who hunt goannas for food consider the perentie to be a high-risk (but tasty) quarry. Debate is growing as to whether goannas are venomous. The incessant bleeding caused by goanna bites had been thought to be the result of
bacterial infection Pathogenic bacteria are bacteria that can cause disease. This article focuses on the bacteria that are pathogenic to humans. Most species of bacteria are harmless and are often beneficial but others can cause infectious diseases. The number of ...
, but a 2005 study suggested monitor lizards (including goannas) are venomous and have oral toxin-producing glands. The goanna's hefty tail can be dangerous when swung, much like a crocodile's tail; small children and dogs have been knocked down by such attacks. Often, victims in goanna attacks are bystanders, watching the person antagonising the goanna. Alarmed goannas can mistake standing humans for trees and attempt to climb them to safety, which is painful and can be distressing for both human and goanna.


Conservation status

Goannas are protected species throughout Australia.


Culture and folklore

Goannas have a prominent place in the culture of indigenous Australians. This includes totemic relationships, anthropomorphic representations in
dreamtime The Dreaming, also referred to as Dreamtime, is a term devised by early anthropologists to refer to a religio-cultural worldview attributed to Australian Aboriginal mythology, Australian Aboriginal beliefs. It was originally used by Francis Ja ...
stories, and as a food source. Representations of goannas are common in indigenous artwork, not just as food, but also as a symbolic spiritual motif. Smaller goannas and the mighty perentie are often considered two different animals when appearing in aboriginal works, as in the story "How the Goanna and Perentie Got Their Colours". European settlers perpetuated several
old wives' tale An old wives' tale is a supposed truth which is actually spurious or a superstition. It can be said sometimes to be a type of urban legend, said to be passed down by older women to a younger generation. Such tales are considered superstition, fol ...
s about goanna habits and abilities; some of these have persisted in modern folklore among campers and bushmen. This includes the above-mentioned exaggeration of goannas dragging off sheep from shepherds' flocks in the night. This might even be exaggerated into child-snatching, rivalling
drop bear The drop bear (sometimes dropbear) is a hoax in contemporary Australian folklore featuring a predatory, carnivorous version of the koala. This imaginary animal is commonly spoken about in tall tales designed to scare tourists. While koalas are t ...
s (attack
koala The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the womb ...
s) as a tourist scarer, but probably more convincing due to the reptiles' carnivorous nature and fearsome appearance. A common tale was that the bite of a goanna was infused with a powerful, incurable
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
. Every year after the bite (or every seven years), the wound would flare up again. For many years, herpetologists generally believed goannas were nonvenomous, and lingering illness from their bites was due solely to
infection An infection is the invasion of tissues by pathogens, their multiplication, and the reaction of host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmissible disease or communicable dise ...
and
septicaemia Sepsis, formerly known as septicemia (septicaemia in British English) or blood poisoning, is a life-threatening condition that arises when the body's response to infection causes injury to its own tissues and organs. This initial stage is follo ...
as a result of their saliva being rife with
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
from carrion and other food sources. However, in 2005, researchers at the
University of Melbourne The University of Melbourne is a public research university located in Melbourne, Australia. Founded in 1853, it is Australia's second oldest university and the oldest in Victoria. Its main campus is located in Parkville, an inner suburb nor ...
announced that oral venom glands had been found in both goannas and iguanas.Goanna venom rocks the reptile record
, UniNews Vol. 14, No. 22 28 November - 12 December 2005, University of Melbourne, Retrieved 8 March 2006 Because the goanna regularly eats snakes (which may involve a fierce struggle), including venomous species, they are often said to be immune to
snake venom Snake venom is a highly toxic saliva containing zootoxins that facilitates in the immobilization and digestion of prey. This also provides defense against threats. Snake venom is injected by unique fangs during a bite, whereas some species are a ...
. However, no evidence found suggests an actual venom
immunity Immunity may refer to: Medicine * Immunity (medical), resistance of an organism to infection or disease * ''Immunity'' (journal), a scientific journal published by Cell Press Biology * Immune system Engineering * Radiofrequence immunity desc ...
. Other stories say that the lizard eats a legendary plant, or drinks from a healing
spring Spring(s) may refer to: Common uses * Spring (season) Spring, also known as springtime, is one of the four temperate seasons, succeeding winter and preceding summer. There are various technical definitions of spring, but local usage of ...
, which neutralises the venom. (This idea is immortalised in
Banjo Paterson Andrew Barton "Banjo" Paterson, (17 February 18645 February 1941) was an Australian bush poet, journalist and author. He wrote many ballads and poems about Australian life, focusing particularly on the rural and outback areas, including the ...
's humorous poem "Johnson's Antidote".) Goanna fat or oil has been anecdotally imbued with mystical healing properties (possibly in connection with their supposed venom immunity). Aboriginal people traditionally used goanna oil as an important bush medicine, and it also became a common medicine among Caucasians shortly after British settlement in Australia. Said to be a cure-all, and possessing amazing powers of penetration (passing through metal as if it were not there), it was sold among early settlers like
snake oil Snake oil is a term used to describe deceptive marketing, health care fraud, or a scam. Similarly, "snake oil salesman" is a common expression used to describe someone who sells, promotes, or is a general proponent of some valueless or fraud ...
in the
Old West The American frontier, also known as the Old West or the Wild West, encompasses the geography, history, folklore, and culture associated with the forward wave of American expansion in mainland North America that began with European colonial ...
of North America. A goanna features as the heroic figure Mr Lizard in the Australian author
May Gibbs Cecilia May Gibbs MBE (17 January 1877 – 27 November 1969) was an Australian children's author, illustrator, and cartoonist. She is best known for her gumnut babies (also known as "bush babies" or "bush fairies"), and the book '' Snugglepot ...
’ children's books ''
Snugglepot and Cuddlepie ''Snugglepot and Cuddlepie'' is a series of books written by Australian author May Gibbs. The books chronicle the adventures of the eponymous Snugglepot and Cuddlepie. The central story arc concerns Snugglepot and Cuddlepie (who are essentially ...
''. A bronze statue of the goanna Mr Lizard has been placed outside the State Library of Victoria. The villain in the Disney film ''
The Rescuers Down Under ''The Rescuers Down Under'' is a 1990 American animated adventure film produced by Walt Disney Feature Animation and released by Walt Disney Pictures. The 29th Disney animated feature film and the second movie to be produced during the Disne ...
'', Percival C. McLeach, has a pet goanna named Joanna.


References


Further reading

* Cogger, H. (1967). ''Australian Reptiles in Colour''. Sydney: A. H. & A. W. Reed, * King, Dennis & Green, Brian. 1999. ''Goannas: The Biology of Varanid Lizards''. University of New South Wales Press. * Underhill, D. (1993). ''Australia's Dangerous Creatures''. Sydney: Reader's Digest.


External links


Johnson's Antidote
by
Banjo Paterson Andrew Barton "Banjo" Paterson, (17 February 18645 February 1941) was an Australian bush poet, journalist and author. He wrote many ballads and poems about Australian life, focusing particularly on the rural and outback areas, including the ...
.
A Run-In With An Australian Desert Lizard
(
NPR National Public Radio (NPR, stylized in all lowercase) is an American privately and state funded nonprofit media organization headquartered in Washington, D.C., with its NPR West headquarters in Culver City, California. It differs from other ...
)
Waitress removes goanna from restaurant
(includes video) {{Varanoidea Monitor lizards of Australia Bushfood Bush medicine Australian Aboriginal bushcraft Reptile common names