Geospiza Fuliginosa
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The small ground finch (''Geospiza fuliginosa'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Thraupidae The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
.
Endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the
Galápagos Islands The Galápagos Islands (Spanish: , , ) are an archipelago of volcanic islands. They are distributed on each side of the equator in the Pacific Ocean, surrounding the centre of the Western Hemisphere, and are part of the Republic of Ecuador ...
, it is common and widespread in
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
, woodland, and other habitats on most islands in the archipelago. It commonly feeds on small seeds and
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s from the skins of Galápagos land and
marine iguana The marine iguana (''Amblyrhynchus cristatus''), also known as the sea iguana, saltwater iguana, or Galápagos marine iguana, is a species of iguana found only on the Galápagos Islands (Ecuador). Unique among modern lizards, it is a marine repti ...
s and
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies (13 ...
s.


Taxonomy and systematics

The small ground finch is one of
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or t ...
, a group of closely related birds which
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation t ...
on the Galápagos Islands. The group is related to the ''
Tiaris The yellow-faced grassquit (''Tiaris olivaceus'') is a passerine bird in the tanager family Thraupidae and is the only member of the genus ''Tiaris''. It is native to the Central America, South America, and the Caribbean. Taxonomy In 1760 the F ...
'' grassquits, which are found in South America and the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
. When
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
first collected the species in 1835, he thought it was a
finch The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
.
John Gould John Gould (; 14 September 1804 – 3 February 1881) was an English ornithologist. He published a number of monographs on birds, illustrated by plates produced by his wife, Elizabeth Gould, and several other artists, including Edward Lear, ...
, who officially described Darwin's specimens, agreed, placing it in the genus ''
Fringilla The genus ''Fringilla'' is a small group of finches from the Old World, which are the only species in the subfamily Fringillinae. The genus name ''Fringilla'' is Latin for "finch". Taxonomy The genus ''Fringilla'' was introduced in 1758 by the ...
'' with the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
finches. By 1841, Gould had changed his mind, moving this and five other species into the new genus ''
Geospiza ''Geospiza'' is a genus of bird in the tanager family Thraupidae. All species in the genus are endemic to the Galápagos Islands. Together with related genera, they are collectively known as Darwin's finches. Although in the past, they were class ...
'' — still a genus of finches, but distinct from those of the Old World. DNA research has now shown that all of Darwin's "finches" are actually
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
s. The name ''Geospiza'' is a combination of the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ''geo''-, meaning "ground-", and ''spiza'', meaning finch. The specific name ''fuliginosa'' is late
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "sooty". It is known to
hybridize Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
(rarely) with the
medium ground finch The medium ground finch (''Geospiza fortis'') is a species of bird in the family Thraupidae. It is endemic to the Galapagos Islands. Its primary natural habitat is tropical shrubland. One of Darwin's finches, the species was the first which sci ...
.


Description

The small ground finch is the smallest of the ground finches, measuring in length. Its
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
is short and pointed, with a slightly curved culmen. On average, its beak is smaller than that of the
medium ground finch The medium ground finch (''Geospiza fortis'') is a species of bird in the family Thraupidae. It is endemic to the Galapagos Islands. Its primary natural habitat is tropical shrubland. One of Darwin's finches, the species was the first which sci ...
, but there is a significant overlap in size between the two, particularly on islands where only one of the two species exists. On islands where the two species compete directly, the difference between their beaks are greater. The male is black with white-tipped undertail coverts, while the female and young are brown with streaked underparts. There are observable phenotypic differences between finches that live in lowlands and ones that live in highlands, and this change is most likely attributed to adaptation. The finches seen in highlands have larger, more pointed beaks and smaller feet and claws compared to the lowland variety. These finches are on a cline (series of biocommunities on a continuous gradient), and individuals in the hybrid zone have intermediate traits. This is an example of parapatric speciation, where the elevation gradient of 560 meters causes differentiation in traits, but hybrids are well adapted in their “hybrid zone.”


Habitat and range

Like all but one of the other Darwin's finches, the small ground finch is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the Galápagos Islands. Abundant and widespread, it is found on every island in the archipelago except for the
Genovesa Genovesa Island ( Spanish: ''Isla Genovesa''), referred to in English as Tower Island, is a shield volcano in the Galápagos Islands in the eastern Pacific Ocean. The island occupies about , and its maximum elevation is . The horse-shoe shaped ...
,
Wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
and Darwin islands. It is most common in arid coastal and transition areas, though it moves into the highlands following the breeding season.


Behavior


Feeding

Like the other Galápagos ground finches, the small ground finch is an
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
with a preference for vegetable matter. It feeds primarily on the ground or in low vegetation, eating seeds, buds, flowers, leaves and the occasional insect. It forms
symbiotic relationship Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
s with
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies (13 ...
s and both
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
and
Galápagos land iguana The Galápagos land iguana (''Conolophus subcristatus'') is a very large species of lizard in the family Iguanidae. It is one of three species of the genus ''Conolophus''. It is endemic to the Galápagos Islands, in the dry lowlands of the island ...
s by
gleaning Gleaning is the act of collecting leftover crops from farmers' fields after they have been commercially harvested or on fields where it is not economically profitable to harvest. It is a practice described in the Hebrew Bible that became a legall ...
parasites from their skins.


Voice

The small ground finch's song is rapid and weak and is transcribed as "''twichooo-twichooo''" or "''teur-weee''".


Conservation and threats

Although the population size of the small ground finch has not been quantified, it is described as common across the Galápagos, and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
lists it as a species of
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
. Its numbers seem to be stable, and neither its population size nor its range size appear to approach thresholds for concern. However, like all endemic wildlife on the Galápagos Islands, it is impacted by some human activities, including fires, overgrazing by domestic and feral animals, and the introduction of exotic species. It is found in ten of the
Important Bird Area An Important Bird and Biodiversity Area (IBA) is an area identified using an internationally agreed set of criteria as being globally important for the conservation of bird populations. IBA was developed and sites are identified by BirdLife Int ...
s established on the islands. The species suffers from high mortality rates from the parasitic fly, ranging from 16% to 95% over a four-year period (2002-2006).O'connor, J, & Robertson, J (2014). Darwin's Finch Begging Intensity Does Not Honestly Signal Need in Parasitised Nests. Ethology, 228, 37.


Note


References


Cited sources

* * {{Taxonbar, from=Q580955 Geospiza Endemic birds of the Galápagos Islands Birds described in 1837 Taxonomy articles created by Polbot