Gymnocanthus Tricuspis
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The Arctic staghorn sculpin (''Gymnocanthus tricuspis'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
belonging to the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...
, the typical sculpins. This sculpin is found in the Arctic Ocean and the northern Atlantic Ocean.


Taxonomy

The Arctic staghorn sculpin was first formally described as ''Cottus tricuspis'' in 1830 by the Danish
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Johan Reinhardt Johannes Christopher Hagemann Reinhardt (23 December 1778 – 31 October 1845), sometimes called J. C. H. Reinhardt, was a professor in zoology at the University of Copenhagen. Born in Rendalen parish in Norway, his father, Johannes Henrik Reinhar ...
with the type locality given as
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''tricuspis'' means "three cusps", a reference to the three cusps on the upper spine on the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
.


Description

The Arctic staghorn sculpin has
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
s supported by between 10 and 12 spines and between 14 and 17 soft rays while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has 15 to 19 soft rays. It has a dark upper body with two blackish brown bands on the flanks being made up of dark blotches. There is an irregular line of dark spots underneath the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
and the underside is pale, yellowish in females, the males having roundish white spots on their bellies. The dorsal fins have light and dark stripes while the
caudal Caudal may refer to: Anatomy * Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism * Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into the ...
, anal and
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s are pale, with a yellow tip and 4-5 diagonal lines of dark spots in the pectoral fins. There are flat, bony plates on the back and the top of the head. The scales of the lateral line scales are tubules embedded in the skin. There are no, or just a few, cirri on the head or body. The uppermost spine on the
preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...
is long and has 1 or more recurved spinules on its upper surface. This species is identified from its congeners by the lack of tubercles to the rear of the eyes and on the nape, a thinner interorbital space which has a width of less than 10% the length of the head and by the possession of many small plates between the eyes. This species has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of .


Distribution and habitat

The Arctic staghorn sculpin is found in teh Arctic Ocean and the northeastern and northwestern Atlantic Ocean. In North American waters it is normally found as far south as the
Gulf of St Lawrence , image = Baie de la Tour.jpg , alt = , caption = Gulf of St. Lawrence from Anticosti National Park, Quebec , image_bathymetry = Golfe Saint-Laurent Depths fr.svg , alt_bathymetry = Bathymetry ...
, occasionally reaching
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
. In the European waters it ranges from the eastern coasts of Greenland, Iceland, along the northern coast of Norway north to the
White Sea The White Sea (russian: Белое море, ''Béloye móre''; Karelian and fi, Vienanmeri, lit. Dvina Sea; yrk, Сэрако ямʼ, ''Serako yam'') is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is su ...
and the
Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian territo ...
to
Svalbard Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
and
Novaya Zemlya Novaya Zemlya (, also , ; rus, Но́вая Земля́, p=ˈnovəjə zʲɪmˈlʲa, ) is an archipelago in northern Russia. It is situated in the Arctic Ocean, in the extreme northeast of Europe, with Cape Flissingsky, on the northern island, ...
. This is a benthic fish which lives in shallow water near to shore down to depths of around , although it is normally dound at depths of less than . It has been recorded on a variety of substrates including mud, gravel and rock, burrowing into sand and muddy-sand substrates.


Biology

The Arctic staghorn sculpin is a predator of
polychaetes Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are mad ...
,
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s,
krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...
, benthic
amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far descr ...
s and echiurids. They in turn are preyed on by larger fishes such as
Arctic cod ''Arctogadus glacialis'', known also with ambiguous common names Arctic cod and polar cod, is an Arctic species of fish in the cod family Gadidae, related to the true cod (genus ''Gadus''). ''Arctogadus glacialis'' is found in icy water. They gr ...
,
Atlantic cod The Atlantic cod (''Gadus morhua'') is a benthopelagic fish of the family Gadidae, widely consumed by humans. It is also commercially known as cod or codling.Bering flounder and polar eelpout. They spawn in the late autumn to winter, the females laying 2,000 to 5,500 demersal eggs which hatch into pelagic larvae and juveniles which descend to the sea bed lengths of . They are sexually mature by 5–6 years old or .


References

{{Taxonbar, from=Q518665 Fish described in 1830 Arctic staghorn sculpin Taxa named by Johan Reinhardt