''Gulella'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of very small air-breathing
land snail
A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
s,
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate
Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is e ...
s in the family
Streptaxidae
Streptaxidae is a family of carnivorous air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade
Stylommatophora. Six Streptaxidae subfamilies are accepted in the 2005 taxonomy of the Gastropoda by Bouchet & Rocroi.
Stre ...
.
[MolluscaBase eds. (2021). MolluscaBase. Gulella L. Pfeiffer, 1856. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=818295 on 2021-01-11]
Until recently, species which are now in the genus ''
Costigulella'' were included here.
Distribution
Distribution of the genus ''Gulella'' include:
*
Afrotropical
The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majority of the Arabian Peninsula, the island of Madagascar, southern Iran and extreme southwestern Pakistan, and the island ...
:
(20 species)
[Wronski T. & Hausdorf B. (2010). "Diversity and body-size patterns of land snails in rain forests in Uganda". '' Journal of Molluscan Studies'' 76(1): 87-100. .]
* Arabia
*
Madagascar
*
Comoros
The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
*
the Seychelles
Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...
*
Mauritius[Sutcharit C., Naggs F., Wade C. M., Fontanilla I. & Panha S. (2010). "The new family Diapheridae, a new species of ''Diaphera'' Albers from Thailand, and the position of the Diapheridae within a molecular phylogeny of the Streptaxoidea (Pulmonata: Stylommatophora)". '' Zoological Journal of the Linnean Society'' 160: 1-16. .]
Species
Species within the large genus ''Gullela'' include:
* ''Gulella aberdarensis''
Preston, 1913
* ''Gulella acutidens''
(O. Boettger, 1905)
* ''Gulella adami''
van Bruggen, 1994
* ''Gulella aequidentata''
(E. A. Smith, 1890)
* ''Gulella afrooccidentalis''
Verdcourt, 2004
* ''Gulella albersi''
(L. Pfeiffer, 1855)
* ''Gulella albinus''
van Bruggen & Van Goethem, 1999
* ''Gulella albogilva''
Germain, 1934
* ''Gulella aliciae''
(Melvill & Ponsonby, 1907)
* ''Gulella alleni''
Verdcourt, 1974
* ''Gulella alutacea''
Connolly, 1932
* ''Gulella amaniensis''
Verdcourt, 1953
* ''Gulella ambalaniranae''
Emberton, 2001
* ''Gulella ambanikelia''
Emberton, 2001
* ''Gulella ambodipelomosiae'' Emberton, 2002
[Emberton K. C. (2002). "''Parvedentulina'' and edentate ''Gulella'' of Madagascar (Gastropoda: Stylommatophora: Streptaxidae)". ''Archiv für Molluskenkunde'' 131(1/2): 67-165]
abstract
* ''Gulella ambrensis''
Emberton, 2001
* ''Gulella amicta''
(E. A. Smith, 1890)
* ''Gulella analamerae''
Emberton, 2001
* ''Gulella andreana''
Fischer-Piette, Blanc, F. & Vukadinovic, 1974
* ''Gulella andriantanteliae''
[Emberton K C., Slapcinsky J., Campbell C. A., Rakotondrazafy J. A., Andriamiarison T. N. & Emberton J. D. (2010). "Terrestrial mollusks of Andriantantely Massif, Eastern Madagascar, with descriptions of 36 new species (Gastropoda: Caenogastropoda; Pulmonata)". ''Archiv für Molluskenkunde'' 139(1): 71-141. ]
* ''Gulella ankaranaensis''
Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
* ''Gulella annibiodiversitatis''
van Bruggen, 2011
* ''Gulella antelmeana''
Peile, 1936
* ''Gulella antongilae''
Emberton, 2001
* ''Gulella appletoni''
van Bruggen, 1975
* ''Gulella aprosdoketa''
Connolly, 1939
* ''Gulella aranearum''
van Bruggen, 1986
* ''Gulella argoudi''
Griffiths, 2000
* ''Gulella arnoldi''
(Sturany, 1898)
* ''Gulella arthuri''
(Dautzenberg, 1890)
* ''Gulella atewana''
de Winter, 1996
* ''Gulella aversostriata''
Verdcourt, 1985
* ''Gulella azeitonae''
D. Holyoak & G. Holyoak, 2020
* ''Gulella baccata''
(Preston, 1913)
* ''Gulella bancoensis''
de Winter, 1996
* ''Gulella barbarae''
Connolly, 1929
* ''Gulella barnardi''
van Bruggen, 1965
* ''Gulella bassetti''
Burnup, 1925
* ''Gulella beandreana''
Emberton, 2001[Emberton K. C. & Griffiths O. L. (2009). "Pulmonate land mollusks from three rainforest transects in northeastern Madagascar: part 1 of 2, with descriptions of 12 new species and one new subspecies (Gastropoda: Pulmonata)". ''Archiv für Molluskenkunde'' 138(2): 137-167. .]
* ''Gulella bebokae''
Emberton, 2001
* ''Gulella bemarahae''
Emberton, 2001
* ''Gulella bemoka''
Emberton, 2001
* ''Gulella benetecta''
Connolly, 1930
* ''Gulella benjamini''
Emberton & Pearce, 2000
* ''Gulella bernardi''
van Bruggen & Van Goethem, 1997
* ''Gulella betamponae''
Emberton, 2002
* ''Gulella bicolor''
(T. Hutton, 1834)
* ''Gulella bitzeense''
Connolly, 1922
* ''Gulella bobaombiae''
Emberton, 2001
* ''Gulella bolocoensis''
Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
* ''Gulella bomolensis''
Verdcourt, 1953
* ''Gulella bomvana''
Cole & Herbert, 2009
* ''Gulella bondi''
Verdcourt, 1962
* ''Gulella bouchardi''
Fischer-Piette, Blanc, F. & Vukadinovic, 1974
* ''Gulella boucheti''
Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
* ''Gulella bowkerae''
(Melvill & Ponsonby, 1892)
* ''Gulella brevicula''
(Morelet, 1882)
* ''Gulella brevis''
(Thiele, 1911)
* ''Gulella bruggeni''
Cole & Herbert, 2009
* ''Gulella burnupi''
(Melvill & Ponsonby, 1897)
* ''Gulella burungaensis''
(Preston, 1913)
* ''Gulella bushmanensis''
Burnup, 1926
* ''Gulella cacosystata''
Germain, 1934
* ''Gulella cacuminata''
Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
* ''Gulella cairnsi''
(Melvill & Ponsonby, 1897)
* ''Gulella callosa''
(Morelet, 1881)
* ''Gulella calopasa''
(Melvill & Ponsonby, 1903)
* ''Gulella camerani''
(Pollonera, 1906)
* ''Gulella cancellata''
Connolly, 1922
* ''Gulella candela''
Connolly, 1922
* ''Gulella candidula''
(Morelet, 1889)
* ''Gulella capmini''
Emberton, 2001
* ''Gulella cara''
Pilsbry, 1919 [Pilsbry H. A. (1919). "A review of the land mollusks of the Belgian Congo chiefly based on the collection of the American Museum Congo Expedition, 1909–1915". ''Bulletin of the American Museum of Natural History'' 40(1): 1-370. page 7, 215-216, 228, 231, 233-234.]
* ''Gulella carea''
(Preston, 1913)
* ''Gulella caroli''
(Kobelt, 1913)
* ''Gulella carpenteri''
Connolly, 1931
* ''Gulella caryatis''
(Melvill & Ponsonby, 1898)
* ''Gulella cavidens''
(E. von Martens, 1876)
* ''Gulella ceciliae''
van Bruggen, 1971
* ''Gulella celestinae''
Emberton, 2001
* ''Gulella cerea''
(Dunker, 1848)
* ''Gulella chapini''
Pilsbry, 1919
* ''Gulella cheranganiensis''
Germain, 1934
* ''Gulella chi''
Burnup, 1926
* ''Gulella claustralis''
Connolly, 1939
* ''Gulella claustrum''
(Preston, 1913)
* ''Gulella coarctata''
(D'Ailly, 1910)
* ''Gulella coarti''
(Dautzenberg & Germain, 1914)
* ''Gulella coeni''
(Preston, 1913)
* ''Gulella collicola''
van Bruggen, 1966
* ''Gulella columella''
(E. A. Smith, 1903)
* ''Gulella columna''
Emberton, 2001
* ''Gulella columnella''
(Melvill & Ponsonby, 1901)
* ''Gulella commoda''
(E. A. Smith, 1903)
* ''Gulella comorensis''
(E. von Martens, 1876)
* ''Gulella conicodentata''
K. L. Pfeiffer, 1952
* ''Gulella connollyi''
(Melvill & Ponsonby, 1909)
* ''Gulella conradti''
(E. von Martens, 1895)
* ''Gulella consanguinea''
(E. A. Smith, 1890)
* ''Gulella consobrina''
(Ancey, 1892)
* ''Gulella consociata''
(E. A. Smith, 1890)
* ''Gulella constricta''
Emberton, 2001
* ''Gulella contingens''
Burnup, 1925
* ''Gulella contraria''
Connolly, 1932
* ''Gulella copiosa''
(Preston, 1913)
* ''Gulella corneola''
(Morelet, 1886)
* ''Gulella costellata''
(Morelet, 1882)
* ''Gulella crassidens''
(L. Pfeiffer, 1859)
* ''Gulella crassilabris''
(Craven, 1881)
* ''Gulella craterodon''
(Melvill & Ponsonby, 1903)
* ''Gulella cruciata''
(E. von Martens, 1900)
* ''Gulella crux''
van Bruggen, 2011
* ''Gulella crystallum''
(Morelet, 1848)
* ''Gulella cupula''
van Bruggen & Van Goethem, 1999
* ''Gulella curvicolumella''
(Preston, 1913)
* ''Gulella curvilamella''
(E. A. Smith, 1890)
* ''Gulella cyclochilus''
Degner, 1934
* ''Gulella cylindrica''
(L. Pfeiffer, 1852)
* ''Gulella daedalea''
(Melvill & Ponsonby, 1903)
* ''Gulella darglensis''
(Melvill & Ponsonby, 1908)
* ''Gulella dartevillei''
Verdcourt, 1962
* ''Gulella dautzenbergi''
Connolly, 1928
* ''Gulella davisae''
Herbert, 2016
* ''Gulella decaryi''
Fischer-Piette & Vukadinovic, 1974
* ''Gulella decussatula''
(Preston, 1913)
* ''Gulella dejae''
Bursey & Herbert, 2004
* ''Gulella delicatula''
(L. Pfeiffer, 1857)
* ''Gulella dentiens''
(Morelet, 1883)
* ''Gulella devia''
Connolly, 1931
* ''Gulella deviae''
Herbert, 2006[Herbert D. (2006). "Rediscovery of the type species of ''Euonyma'' (Subulinidae) and observations on South African species of ''Gulella'' (Streptaxidae), with description of two new species (Gastropoda: Eupulmonata)". ''Journal of Natural History'' 40(17-18): 1063-1081. .]
* ''Gulella dextrorsa''
Burnup, 1925
* ''Gulella diabensis''
Connolly, 1939
* ''Gulella diodon''
(Morelet, 1882)
* ''Gulella discrepans''
(Sturany, 1898)
* ''Gulella disseminata''
(Preston, 1913)
* ''Gulella distincta''
(Melvill & Ponsonby, 1893)
* ''Gulella dohrni''
(E. A. Smith, 1882)
* ''Gulella dolichos''
Verdcourt, 1985
* ''Gulella doliolum''
(Morelet, 1867)
* ''Gulella dorri''
(Dautzenberg, 1890)
* ''Gulella drakensbergensis''
(Melvill & Ponsonby, 1893)
* ''Gulella duncani''
Connolly, 1930
* ''Gulella dunkeri''
(L. Pfeiffer, 1856)
* ''Gulella dupuisi''
Connolly, 1922
* ''Gulella dupuyana''
(Crosse, 1876)
* ''Gulella ectodentata''
van Bruggen & Van Goethem, 1999
* ''Gulella elliptica''
(Melvill & Ponsonby, 1898)
* ''Gulella enneodon''
Connolly, 1922
* ''Gulella eoryi''
Verdcourt, 2004
* ''Gulella erugo''
Rowson, Foster, Seddon & Tattersfield, 2019
* ''Gulella escalarai''
Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
* ''Gulella eussoensis''
(Preston, 1913)
* ''Gulella euthymia''
(Melvill & Ponsonby, 1893)
* ''Gulella excavata''
(E. von Martens, 1892)
* ''Gulella excruciata''
Connolly, 1931
* ''Gulella exogonia''
(E. von Martens, 1895)
* ''Gulella expatriata''
(Preston, 1910)
* ''Gulella falconi''
Burnup, 1925
* ''Gulella farquhari''
(Melvill & Ponsonby, 1895)
* ''Gulella fernandensis''
Ortiz de Zárate López & Ortiz de Zárate Rocandio, 1956
* ''Gulella filix''
Connolly, 1922
* ''Gulella fischerpiettei''
Emberton, 2001
* ''Gulella formosa''
(Melvill & Ponsonby, 1898)
* ''Gulella fortidentata''
(E.A. Smith, 1890)
* ''Gulella fotobohitrae''
Emberton, 2001
* ''Gulella foveolata''
Preston, 1913
* ''Gulella framesi''
Burnup, 1926
* ''Gulella fraudator''
Connolly, 1939
* ''Gulella funerea''
(Preston, 1913)
* ''Gulella galactochila''
(Crosse, 1885)
* ''Gulella gallorum''
Fischer-Piette, Blanc, F. & Salvat, 1975
* ''Gulella garambae''
van Bruggen & Van Goethem, 1999
* ''Gulella genialis''
(Melvill & Ponsonby, 1903)
* ''Gulella gestroi''
(Germain, 1915)
* ''Gulella girardi''
(Kobelt, 1904)
* ''Gulella glabra''
(Morelet, 1882)
* ''Gulella godfreyi''
Burnup, 1925
* ''Gulella gouldi''
(L. Pfeiffer, 1856)
* ''Gulella greenwayi''
Verdcourt, 1953
* ''Gulella griffithsi''
Emberton, 2001
* ''Gulella guilielmi''
van Bruggen & Van Goethem, 1998
* ''Gulella gwendolinae''
(Preston, 1910)
* ''Gulella hadroglossa''
Herbert, 2016
* ''Gulella hafa''
Emberton, 2001
* ''Gulella hafahafa''
Emberton, 2001
* ''Gulella hamerae''
Bursey & Herbert, 2004
* ''Gulella handeiensis''
Verdcourt, 1953
* ''Gulella harriesi''
Burnup, 1926
* ''Gulella haullevillei''
(Dautzenberg & Germain, 1914)
* ''Gulella hector''
(Preston, 1913)
* ''Gulella helichrysophila''
Germain, 1934
* ''Gulella hemmingi''
Verdcourt, 1963
* ''Gulella herberti''
van Bruggen, 2004
* ''Gulella heteromphala''
Pilsbry, 1919
* ''Gulella hildae''
van Bruggen, 2001
* ''Gulella himerothales''
(Melvill & Ponsonby, 1903)
* ''Gulella hoqensis''
Neubert, 2004
* ''Gulella hordeum''
(Morelet, 1879)
* ''Gulella hughscotti''
Verdcourt, 1980
* ''Gulella humbloti''
(Morelet, 1886)
* ''Gulella impedita''
Connolly, 1922
* ''Gulella inconspicua''
(Thiele, 1911)
* ''Gulella incurvidens''
van Bruggen, 1972
* ''Gulella infans''
(Craven, 1881)
* ''Gulella infrendens''
(E. von Martens, 1866)
* ''Gulella ingloria''
(Preston, 1913)
* ''Gulella inhluzaniensis''
(Burnup, 1914)
* ''Gulella inobstructa''
van Bruggen, 1965
* ''Gulella insolita''
(E. A. Smith, 1903)
* ''Gulella instabilis''
(Sturany, 1898)
* ''Gulella insularis''
(Girard, 1894)
* ''Gulella insulincola''
van Bruggen, 1975
* ''Gulella intradentata''
(Preston, 1913)
* ''Gulella intrusa''
Verdcourt, 1956
* ''Gulella io''
Verdcourt, 1974
* ''Gulella iridescens''
(Preston, 1913)
* ''Gulella isipingoensis''
(Sturany, 1898)
* ''Gulella isseli''
(Paladilhe, 1872)
* ''Gulella jaominai''
Emberton, 2001
* ''Gulella jod''
(Preston, 1910)
* ''Gulella johannae''
van Bruggen, 2006
* ''Gulella johannesburgensis''
(Melvill & Ponsonby, 1907)
* ''Gulella jongkindi''
de Winter, 1996
* ''Gulella josephinae''
Emberton, 2001
* ''Gulella juxtidens''
(Melvill & Ponsonby, 1899)
* ''Gulella kelibea''
Emberton, 2001
* ''Gulella kelimolotra''
Emberton, 2002
* ''Gulella kendrae''
Emberton & Griffiths, 2009
* ''Gulella kidundae''
Adam, 1965
* ''Gulella kigeziensis''
(Preston, 1913)
* ''Gulella kohllarseni''
(Haas, 1936)
* ''Gulella kosiensis''
(Melvill & Ponsonby, 1908)
* ''Gulella kraussi''
(L. Pfeiffer, 1856)
* ''Gulella kulalensis''
Verdcourt, 1962
* ''Gulella labiotuberculata''
Connolly, 1942
* ''Gulella lacuna''
(Preston, 1911)
* ''Gulella laevigata''
(Dohrn, 1865)
* ''Gulella laevorsa''
Burnup, 1925
* ''Gulella lambda''
Degner, 1934
* ''Gulella lamyi''
(Dautzenberg & Germain, 1914)
* ''Gulella landianiensis''
(Dautzenberg, 1908)
* ''Gulella langei''
Rowson, Foster, Seddon & Tattersfield, 2019
* ''Gulella laninifia''
Emberton, 2002
* ''Gulella laqueus''
(Preston, 1913)
* ''Gulella larva''
(Morelet, 1877)
* ''Gulella latimerae''
Bursey & Herbert, 2004
* ''Gulella lawrencei''
van Bruggen, 1964
* † ''Gulella leakeyi''
Verdcourt, 1963
* ''Gulella lendix''
(E. A. Smith, 1890)
* ''Gulella lessensis''
Pilsbry, 1919
* ''Gulella leucocion''
Connolly, 1929
* ''Gulella lievrouwi''
van Bruggen & Van Goethem, 1999
* ''Gulella lima''
(Preston, 1913)
* ''Gulella lincolni''
Emberton & Griffiths, 2009
* ''Gulella lindae''
Herbert, 2006
* ''Gulella linguidens''
Connolly, 1939
* ''Gulella lissophanes''
(Melvill & Ponsonby, 1892)
* ''Gulella lohabea''
Emberton, 2001
* ''Gulella lornae''
Verdcourt, 1952
* ''Gulella loveridgei''
van Bruggen, 1996
* ''Gulella lubeti''
Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
* ''Gulella lubrica''
(Morelet, 1881)
* ''Gulella ludwigi''
Verdcourt & Venmans, 1953
* ''Gulella magnifica''
Emberton, 2001
* ''Gulella magnolia''
(Connolly, 1912)
* ''Gulella magnorchida''
Emberton, 2001
* ''Gulella mahafinaratra''
Emberton, 2001
* ''Gulella mahagaga''
Emberton, 2001
* ''Gulella mahia''
Emberton, 2001
* ''Gulella malasangiensis''
(Preston, 1913)
* ''Gulella mamellensis''
Griffiths, 2000
* ''Gulella manomboae''
Emberton, 2001
* ''Gulella mariae''
(Melvill & Ponsonby, 1892)
* ''Gulella marionae''
(Preston, 1910)
* ''Gulella maritzburgensis''
(Melvill & Ponsonby, 1893)
* ''Gulella marojejyae''
Emberton, 2001
* ''Gulella masoalae''
Emberton, 2001
* ''Gulella masis iensis''
Pilsbry, 1919
* ''Gulella matavymolotra''
Emberton, 2002
* ''Gulella matumbiensis''
Verdcourt, 1993
* ''Gulella mayottensis''
Connolly, 1925
* † ''Gulella mbili''
Pickford, 2009
* ''Gulella megapex''
Neubert, 2004
* ''Gulella melvilli''
(Burnup, 1914)
* ''Gulella meneleki''
(Preston, 1910)
* ''Gulella menkeana''
(L. Pfeiffer, 1853)
* ''Gulella menkhorsti''
van Bruggen, 2011
* ''Gulella meruensis''
(D'Ailly, 1910)
* ''Gulella mfongosiensis''
Burnup, 1925
* ''Gulella miaranoniae''
Emberton, 2001
* ''Gulella miaryi''
Fischer-Piette & Bedoucha, 1964
* ''Gulella micans''
K. L. Pfeiffer, 1952
* ''Gulella michellae''
Emberton, 2001
* ''Gulella microdina''
(Morelet, 1883)
* ''Gulella microdon''
(Morelet, 1860)
* ''Gulella microrutshuruensis''
van Bruggen, 1995
* ''Gulella microstriata''
Emberton, 2001
* ''Gulella mihomehia''
Emberton, 2001
* ''Gulella mikenoensis''
(Preston, 1913)
* ''Gulella miniata''
(F. Krauss, 1848)
* ''Gulella minuscula''
Emberton & Pearce, 2000
* ''Gulella minuta''
(Morelet, 1889)
* ''Gulella minutissima''
(Thiele, 1911)
* ''Gulella mitsikia''
Emberton, 2001
* ''Gulella mkuu'' R
owson, Seddon & Tattersfield, 2009
* ''Gulella modioliformis''
(Morelet, 1877)
* † ''Gulella moja''
Pickford, 2009
* ''Gulella mongolae''
Ortiz de Zárate Lopez & Ortiz de Zárate Rocandio, 1956
* ''Gulella monodon''
(Morelet, 1873)
* ''Gulella montium''
(D'Ailly, 1910)
* ''Gulella mooiensis''
(Burnup, 1914)
* ''Gulella msoalae''
Emberton, 2001
* ''Gulella multidentata''
(Sturany, 1898)
* ''Gulella munita''
(Melvill & Ponsonby, 1892)
* ''Gulella nakamaroa''
Emberton, 2001
* ''Gulella namorokae''
Emberton, 2001
* ''Gulella natalensis''
(Craven, 1881)
* ''Gulella ndibo''
Cole & Herbert, 2009
* ''Gulella nemoralis''
(Germain, 1915)
* ''Gulella nepia''
Connolly, 1925
* ''Gulella newmani''
Bursey & Herbert, 2004
* ''Gulella newtoni''
(E. A. Smith, 1890)
* ''Gulella ngorogoroensis''
Verdcourt, 1985
* ''Gulella nifikelia''
Emberton, 2001
* † ''Gulella nne''
Pickford, 2009
* ''Gulella noltei''
(Boettger, 1898)
* ''Gulella nosybei''
Emberton, 2001
* ''Gulella nuchalis''
van Bruggen, 2011
* ''Gulella nyikaensis''
(Preston, 1913)
* ''Gulella nyiroensis''
(Preston, 1913)
* ''Gulella obani''
Oke, 2007
* ''Gulella obovata''
(L. Pfeiffer, 1855)
* ''Gulella obstructa''
van Bruggen, 1965
* ''Gulella odhneriana''
Dupuis, 1923
* ''Gulella odietei''
Oke, 2013
* ''Gulella ogbeifuni''
Oke, 2013
* ''Gulella olkokolae''
W. Adam, 1965
* ''Gulella opoboensis''
(Preston, 1914)
* ''Gulella orchida''
Emberton, 2001
* ''Gulella orientalis''
Connolly, 1929
* ''Gulella oryza''
(Morelet, 1882)
* ''Gulella ovalis''
(Thiele, 1911)
* ''Gulella ovularis''
(Morelet, 1886)
* ''Gulella pangumana''
Connolly, 1928
* ''Gulella papyracea''
(Preston, 1913)
* ''Gulella paucidens''
Verdcourt, 1953
* ''Gulella pearcei''
Emberton, 2001
* ''Gulella peculiaris''
(E. A. Smith, 1890)
* ''Gulella penningtoni''
Burnup, 1925
* ''Gulella pentheri''
(Sturany, 1898)
* ''Gulella pentodon''
(Morelet, 1889)
* ''Gulella percivali''
(Preston, 1913)
* ''Gulella pergrata''
(Preston, 1913)
* ''Gulella perissodonta''
(Sturany, 1898)
* ''Gulella perlata''
Connolly, 1922
* ''Gulella perplexa''
Connolly, 1939
* ''Gulella perspicua''
(Melvill & Ponsonby, 1893)
* ''Gulella perspicuaeformis''
(Sturany, 1898)
* ''Gulella pervitrea''
(Preston, 1913)
* ''Gulella petitboucheti''
Emberton, 2001
* ''Gulella pfeifferi''
(Krauss in Kuster, 1844)
* ''Gulella phanerodon''
(Morelet, 1888)
* ''Gulella phragma''
(Melvill & Ponsonby, 1907)
* ''Gulella phyllisae''
Burnup, 1925
* ''Gulella planidens''
(E. von Martens, 1892)
* ''Gulella planti''
(L. Pfeiffer, 1856) - Plant's gulella snail
* ''Gulella platycostata''
Neubert, 2004
* ''Gulella plicigera''
(Morelet, 1886)
* ''Gulella polita''
(Melvill & Ponsonby, 1893)
* ''Gulella polloneriana''
Pilsbry, 1919
* ''Gulella pondoensis''
Connolly, 1939
* ''Gulella ponsonbyi''
(Burnup, 1914)
* ''Gulella porcina''
Connolly, 1930
* ''Gulella poutrini''
(Germain, 1918)
* ''Gulella praelonga''
Connolly, 1922
* ''Gulella premnodes''
(Sturany, 1902)
* ''Gulella prestoni''
Connolly, 1922
* ''Gulella pretoriana''
Connolly, 1932
* ''Gulella princei''
(Preston, 1911)
* ''Gulella protruda''
Neubert & Frank, 1996
* ''Gulella proxima''
van Bruggen, 2014
* ''Gulella pseudandreana''
Emberton, 2001
* ''Gulella pseudolkokolae''
Verdcourt, 2004
* ''Gulella puella''
Connolly, 1929
* ''Gulella pulchella''
(Melvill & Ponsonby, 1893)
* ''Gulella pusilla''
(Morelet, 1881)
* ''Gulella puzeyi''
Connolly, 1939
* ''Gulella queketti''
(Melvill & Ponsonby, 1896)
* ''Gulella raffrayi''
(Bourguignat, 1883)
* ''Gulella rakotoarisoni''
Emberton, 2001
* ''Gulella ranomasina''
Emberton, 2001
* ''Gulella razafyi''
Emberton, 2001
* ''Gulella rectangularis''
(Preston, 1913)
* ''Gulella reeae''
Emberton & Pearce, 2000
* ''Gulella reesi''
(Preston, 1914)
* ''Gulella reniformis''
(Preston, 1913)
* ''Gulella rewgidensis''
Neubert, 2004
* ''Gulella rhodesiana''
(Connolly, 1912)
* ''Gulella roberti''
(Preston, 1910)
* ''Gulella rogersi''
(Melvill & Ponsonby, 1898)
* ''Gulella rubinsterni''
Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
* ''Gulella rugosa''
Emberton, 2001
* ''Gulella rumpiana''
Connolly, 1932
* ''Gulella runa''
Rowson, Foster, Seddon & Tattersfield, 2019
* ''Gulella ruthae''
Emberton, 2002
* ''Gulella rutshuruensis''
Pilsbry, 1919
* ''Gulella ruwenzoriensis'' van Bruggen & van Goethem, 1999
* ''Gulella sahia''
Emberton, 2002
* ''Gulella salpinx''
Herbert, 2002[Herbert D. G. (2002). "''Gulella salpinx'' sp. n., a new critical endangered holoendemic species from the limestone deposits of the Marble Delta, KwaZuluNatal, South Africa (Mollusca: Gastropoda: Streptaxidae)". ''African Invertebrates'' 43: 125-138]
abstract
* ''Gulella sandersoni''
Connolly, 1942
* ''Gulella sankuruensis''
Dartevelle-Puissant, 1936
* ''Gulella satisfacta''
Fischer-Piette, Blanc, C.P., Blanc, F. & Salvat, 1994
* ''Gulella schweinfurthi'' (Thiele, 1910)
* ''Gulella sellae''
(Pollonera, 1906)
* ''Gulella seneciophila''
Germain, 1934
* ''Gulella separata''
(Sturany, 1898)
* ''Gulella sesamum''
(Morelet, 1883)
* ''Gulella sexdentata''
(E. von Martens, 1869)
* ''Gulella shabae''
Adam & Van Goethem, 1978
* ''Gulella shoaensis''
Verdcourt, 1985
* ''Gulella sibasana''
Connolly, 1922
* ''Gulella simplicima''
(Preston, 1911)
* † ''Gulella sita''
Pickford, 2009
* ''Gulella socialis''
Pilsbry, 1919
* ''Gulella socotrensis''
Neubert, 2004
* ''Gulella soror''
(E. A. Smith, 1890)
* ''Gulella soulaiana''
Fischer-Piette, 1973
* ''Gulella spreta''
(Morelet, 1883)
* ''Gulella stolidodea''
Degner, 1934
* ''Gulella streptostelopsis''
Adolph Cornelis van Bruggen, Bruggen, 2007[Adolph Cornelis van Bruggen, van Bruggen A. C. (June 2007). "Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malaŵi 9. Description of ''Gulella streptostelopsis'', a new ''Streptostele''-like species of ''Gulella''". ''Zoologische Mededelingen'' 81(1). http://www.zoologischemededelingen.nl/81/nr01/a01 ]
* ''Gulella strictilabris''
(Ancey, 1898)
* ''Gulella stylodon''
(E. von Martens, 1876)
* ''Gulella suavissima''
(Preston, 1913)
* ''Gulella subflavescens''
(E. A. Smith, 1890)
* ''Gulella subframesi''
Connolly, 1929
* ''Gulella subhyalina''
(E. A. Smith, 1890)
* ''Gulella subkraussi''
Connolly, 1932
* ''Gulella subringens''
(Crosse, 1886)
* ''Gulella subsellae''
Haas, 1936
* ''Gulella sursum''
van Bruggen, 2001
* ''Gulella swaziensis''
Connolly, 1932
* ''Gulella sylvia''
(Melvill & Ponsonby, 1903)
* ''Gulella syngenes''
(Preston, 1913)
* ''Gulella systemanaturae''
van Bruggen, 2008[Adolph Cornelis van Bruggen, Bruggen A. C. van (2008). "Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malaŵi 10. Description of ''Gulella systemanaturae'', a new species from Dedza Mountain". ''Zoologische Mededelingen'' Leiden, 82(2) pp. 9-14]
HTM
PDF
* ''Gulella taitensis''
Verdcourt, 1963
* † ''Gulella tano''
Pickford, 2009
* ''Gulella taolantehezana''
Emberton, 2002
* † ''Gulella tatu''
Pickford, 2009
* ''Gulella tendronia''
Emberton, 2001
* ''Gulella tharfieldensis''
(Melvill & Ponsonby, 1893)
* ''Gulella thompsoni''
K. C. Emberton, Slapcinsky, Campbell, Rakotondrazafy, Andriamiarison & J. D. Emberton, 2010
* ''Gulella tietzae''
Cole & Herbert, 2009
* ''Gulella titania''
Connolly, 1928
* ''Gulella tomlini''
(Preston, 1911)
* ''Gulella tracheia''
Rowson, 2007
* ''Gulella translucida''
K. L. Pfeiffer, 1952
* ''Gulella transnominata''
Connolly, 1925
* ''Gulella triglochis''
(Melvill & Ponsonby, 1903)
* ''Gulella trigona''
(Morelet, 1881)
* ''Gulella triplicina''
(E. von Martens, 1895)
* ''Gulella tripodium''
Connolly, 1939
* ''Gulella tristaoensis''
Connolly, 1922
* ''Gulella tsara''
Emberton, 2001
* ''Gulella tsaratananae''
Emberton, 2001
* ''Gulella tudes''
(E. von Martens, 1895)
* ''Gulella udzungwensis''
van Bruggen, 2003
* ''Gulella ugandensis''
(E. A. Smith, 1901)
* ''Gulella uluguruensis''
Verdcourt, 1962
* ''Gulella umzimvubuensis''
Burnup, 1925
* ''Gulella unidentata''
K. L. Pfeiffer, 1952
* ''Gulella vakinifia''
Emberton, 2001
* ''Gulella vallaris''
(Melvill & Ponsonby, 1907)
* ''Gulella varians''
(E. A. Smith, 1899)
* ''Gulella vatosoa''
Emberton, 2001
* ''Gulella vavakelia''
Emberton, 2001
* ''Gulella ventricosa''
Neubert, 2004
* ''Gulella verdcourti''
van Bruggen, 1966
* ''Gulella vermis''
(Morelet, 1881)
* ''Gulella viae''
Burnup, 1925
* ''Gulella viatoris''
(Preston, 1913)
* ''Gulella vicina''
(E.A. Smith, 1899)
* ''Gulella virungae''
van Bruggen & van Goethem, 1999
* ''Gulella vohimarae''
Emberton, 2001
* ''Gulella vriesiana''
(C. M. F. Ancey, 1885)
* ''Gulella wahlbergi''
(F. Krauss, 1848)
* ''Gulella warrenii''
(Melvill & Ponsonby, 1903)
* ''Gulella wendalinae''
van Bruggen, 1975
* ''Gulella wilmattae''
(Pilsbry & Cockerell, 1933)
* ''Gulella woodhousei''
(Preston, 1913)
* ''Gulella xysila''
(Melvill & Ponsonby, 1907)
* ''Gulella zanaharyi''
Emberton, 2001
* ''Gulella zelota''
(Melvill & Ponsonby, 1907)
* ''Gulella zemenensis''
Verdcourt, 1990
* ''Gulella zuluensis''
Connolly, 1932
* ''Gulella'' sp. 1 sensu Emberton & Griffiths (2009)
* ''Gulella'' sp. 2 sensu Emberton & Griffiths (2009)
* ''Gulella'' sp. 3 sensu Emberton & Griffiths (2009)
synonyms:
* ''Gulella amboniensis'' : synonym of ''Juventigulella amboniensis''
(Tattersfield, 1998) (original combination)
* ''Gulella avakubiensis''
Pilsbry, 1919:
synonym of ''Avakubia avakubiensis''
(Pilsbry, 1919) (superseded combination)
* ''Gulella caryatis diabensis''
Connolly, 1939:
synonym of ''Gulella diabensis''
Connolly, 1939 (original combination)
* ''Gulella conospira''
(Martens, 1892):
synonym of ''Conogulella conospira''
(E. von Martens, 1892) (superseded combination)
* ''Gulella cuspidata''
Verdcourt, 1962: synonym of ''Dadagulella cuspidata''
(Verdcourt, 1962) (original combination)
* ''Gulella foliifera''
(E. von Martens, 1895): synonym of ''Primigulella foliifera''
(E. von Martens, 1895) (superseded combination)
* ''Gulella grossa''
(E. von Martens, 1892): synonym of ''Primigulella grossa''
(E. von Martens, 1892) (superseded combination)
* ''Gulella kuiperi''
de Winter, 2007:
[de Winter A. J. (2007). "''Gulella'' (''Silvigulella'') ''kuiperi'' spec. nov., a new minute land snail from Cameroon (Gastropoda, Pulmonata, Streptaxidae)". ''Basteria'' suppl 3: 69-72.] synonym of ''Silvigulella kuiperi''
(De Winter, 2006) (basionym)
* ''Gulella linguifera''
(Martens, 1895):
synonym of ''Primigulella linguifera''
(E. von Martens, 1895) (superseded combination)
* ''Gulella ndamanyiluensis''
Venmans, 1956: synonym of ''Primigulella ndamanyiluensis''
(Venmans, 1956) (original combination)
* ''Gulella osborni''
Pilsbry, 1919:
synonym of ''Silvigulella osborni''
(Pilsbry, 1919) (original combination)
* ''Gulella pupa''
(Thiele, 1911):
synonym of ''Pupigulella pupa''
(Thiele, 1911) (superseded combination)
* ''Gulella selene''
van Bruggen & van Goethem, 1999:
synonym of ''Dadagulella selene''
(van Bruggen & Van Goethem, 1999) (original combination)
* ''Gulella thomasseti'' is a synonym for ''Glabrennea thomasseti'' (Sykes, 1909)
* ''Gulella usambarica''
K. L. Pfeiffer, 1952: synonym of ''Gulella ludwigi''
Verdcourt & Venmans, 1953 (junior secondary homonym of ''Gulella usambarica''
(Craven, 1880); ''Gulella ludwigi''
Verdcourt & Venmans, 1953 is a replacement name)
* ''Gulella usambarica''
(Craven, 1880): synonym of ''Primigulella usambarica''
(Craven, 1880) (superseded combination)
References
* Bank, R. A. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16th, 2017
Further reading
* Connolly, M. (1922). Notes on African non-marine Mollusca, with descriptions of many new species. The Annals and Magazine of Natural History, (9) 10 (59): 485-517, pl. 14. London.
* Henry Clifden Burnup, Burnup H. C. (1925). "On some South African Gulellae, with descriptions of new species and varieties. Part 1." ''Annals of the Natal Museum'' 5: 101-158; pls. 8-9.
* Pilsbry, H.A. & Cockerell, T.D.A. (1933). African Mollusca, chiefly from the Belgian Congo. Proceedings of the Zoological Society of London 1933: 365–375.
* Verdcourt B. (1985). "New taxa of ''Gulella'' L. Pfr. and ''Ptychotrema'' Mörch (Mollusca, Streptaxidae) from Eastern Africa". ''Journal of Conchology'' 32(1): 109-122
abstract
* Rowson B. & Herbert D.G. (2016). The type species and circumscription of the species-rich Afrotropical snail genus Gulella L. Pfeiffer, 1856, based on anatomical and mtDNA data (Mollusca: Eupulmonata: Streptaxidae). Archiv für Molluskenkunde. 145(1): 69-84. page(s): 76
External links
*
Pfeiffer, L. (1856). Versuch einer Anordnung derr Heliceen nach natürlichen Gruppen. Malakozoologische Blätter. 2: 112-185.Pilsbry, H.A. (1919). A review of the land mollusks of the Belgian Congo chiefly based on the collections of the American Museum Congo Expedition, 1909-1915. Bulletin of the American Museum of Natural History, 40: 1-370, pls I-XXIII
{{Taxonbar, from=Q5617359
Gulella,
Streptaxidae
Taxonomy articles created by Polbot
Taxa named by Ludwig Karl Georg Pfeiffer