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Guava () is a common
tropical fruit A tropical fruit one that typically grows in warm climates, or equatorial areas. Tropical fruits Varieties of tropical fruit include: * Acerola ( West Indian Cherry or Barbados Cherry) *Ackee *Banana *Barbadine (granadilla; maracujá-açu in ...
cultivated in many tropical and subtropical regions. The common guava ''
Psidium guajava ''Psidium guajava'', the common guava, yellow guava, lemon guava, or apple guava is an evergreen shrub or small tree native to the Caribbean, Central America and South America. It is easily pollinated by insects; when cultivated, it is pollina ...
'' (lemon guava, apple guava) is a small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
in the myrtle
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
(
Myrtaceae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All speci ...
), native to
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
and northern South America. The name guava is also given to some other species in the genus ''
Psidium ''Psidium'' is a genus of trees and shrubs in the family Myrtaceae. It is native to warmer parts of the Western Hemisphere (Mexico, Central and South America, the West Indies the Galápagos islands). Taxonomy This genus was described first by ...
'' such as strawberry guava (''
Psidium cattleyanum ''Psidium cattleyanum (World Plants : Psidium cattleianum)'', commonly known as Cattley guava, strawberry guava or cherry guava, is a small tree (2–6 m tall) in the Myrtaceae (myrtle) family. The species is named in honour of English hor ...
'') and to the pineapple guava, ''
Feijoa sellowiana ''Feijoa sellowiana'' is a species of flowering plant in the myrtle family, Myrtaceae. It is native to the highlands of southern Brazil, eastern Paraguay, Uruguay, northern Argentina, and Colombia. It is widely cultivated as an ornamental tree ...
''. In 2019, 55 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton ( United State ...
s of guavas were produced worldwide, led by
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
with 45% of the total. Botanically, guavas are
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
.


Types

The most frequently eaten species, and the one often simply referred to as "the guava", is the apple guava (''
Psidium guayava ''Psidium guajava'', the common guava, yellow guava, lemon guava, or apple guava is an evergreen shrub or small tree native to the Caribbean, Central America and South America. It is easily pollinated by insects; when cultivated, it is pollina ...
''). Guavas are typical
Myrtoideae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All spe ...
, with tough dark heavy leaves that are opposite, simple, elliptic to ovate, and long. The flowers are white, with five petals and numerous
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s. The fruits are many-seeded
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
.


Etymology

The term ''guava'' appears to have been in use since the mid-
16th century The 16th century begins with the Julian year 1501 ( MDI) and ends with either the Julian or the Gregorian year 1600 ( MDC) (depending on the reckoning used; the Gregorian calendar introduced a lapse of 10 days in October 1582). The 16th cent ...
. The name derived from the
Taíno The Taíno were a historic Indigenous peoples of the Caribbean, indigenous people of the Caribbean whose culture has been continued today by Taíno descendant communities and Taíno revivalist communities. At the time of European contact in the ...
, a language of the Awawaks as for ''guava tree'' via the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...
for . It has been adapted in many European and Asian languages, having a similar form.


Origin and distribution

Guavas originated from an area thought to extend from Mexico, Central America or northern South America throughout the Caribbean region. Archaeological sites in Peru yielded evidence of guava cultivation as early as 2500 BCE. Guava was adopted as a crop in subtropical and tropical Asia, parts of the United States (from Tennessee and North Carolina, southward, as well as the west and Hawaii), tropical Africa, South Asia, Southeast Asia, and Oceania. Guavas were introduced to
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, US in the 19th century and are grown there as far north as
Sarasota Sarasota () is a city in Sarasota County on the Gulf Coast of the U.S. state of Florida. The area is renowned for its cultural and environmental amenities, beaches, resorts, and the Sarasota School of Architecture. The city is located in the sout ...
, Chipley,
Waldo Waldo may refer to: People * Waldo (given name), a list of people and fictional characters * Waldo (surname), a list of people * Waldo (footballer) (1934-2019), full name Waldo Machado da Silva, Brazilian footballer Places Canada * Waldo, ...
and
Fort Pierce Fort Pierce is a city in and the county seat of St. Lucie County, Florida, United States. The city is part of the Treasure Coast region of Atlantic Coast Florida. It is also known as the Sunrise City, sister to San Francisco, California, the Suns ...
. However, they are a primary host of the Caribbean fruit fly and must be protected against infestation in areas of Florida where this pest is present. Guavas are cultivated in many tropical and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
countries. Several
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are grown commercially; apple guava and its
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s are those most commonly traded internationally. Guavas also grow in southwestern Europe, specifically the
Costa del Sol The Costa del Sol (literally "Coast of the Sun" or "Sun Coast") is a region in the south of Spain in the autonomous community of Andalusia, comprising the coastal towns and communities along the coastline of the Province of Málaga and the easte ...
on
Málaga Málaga (, ) is a municipality of Spain, capital of the Province of Málaga, in the autonomous community of Andalusia. With a population of 578,460 in 2020, it is the second-most populous city in Andalusia after Seville and the sixth most pop ...
, (Spain) and Greece where guavas have been commercially grown since the middle of the 20th century and they proliferate as cultivars. Mature trees of most species are fairly cold-hardy and can survive temperatures slightly colder than for short periods of time, but younger plants will likely freeze to the ground. Guavas are of interest to home growers in subtropical areas as one of the few tropical fruits that can grow to fruiting size in pots indoors. When grown from seed, guava trees can bear fruit in two years, and can continue to do so for forty years.


Ecology

''Psidium'' species are eaten by the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
, mainly
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s like the Ello Sphinx (''
Erinnyis ello ''Erinnyis ello'', the ello sphinx, is a moth of the family Sphingidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It is distributed from Argentina through Central America to the United Stat ...
''), ''
Eupseudosoma aberrans ''Eupseudosoma aberrans'' is a moth of the family Erebidae first described by William Schaus in 1905. It is found in Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North Amer ...
'', '' E. involutum'', and ''
Hypercompe icasia ''Hypercompe icasia'' is a moth of the family Erebidae first described by Pieter Cramer in 1777. It is widely distributed in South America and is also found on Martinique, Guadeloupe, Saint Martin, Dominica, St. Kitts, Nevis, St. Thomas and P ...
''.
Mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...
s, like '' Pronematus pruni'' and '' Tydeus munsteri'', are known to be crop
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
of the
apple guava ''Psidium guajava'', the common guava, yellow guava, lemon guava, or apple guava is an evergreen shrub or small tree native to the Caribbean, Central America and South America. It is easily pollinated by insects; when cultivated, it is pollina ...
('' P. guajava'') and perhaps other species. The
bacterium Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
''
Erwinia psidii ''Erwinia psidii'' is a Gram-negative bacterium and a phytopathogen of the common guava (''Psidium guajava''), causing rot in branches, flowers and fruits. Recently, it was demonstrated that this species produces two acyl homoserine lactones (' ...
'' causes rot diseases of the apple guava. The fruit is cultivated and favored by humans, and many other animals such as birds consume it, readily dispersing the seeds in their droppings. In
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
, strawberry guava ('' P. littorale'') has become an aggressive
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
threatening
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
to more than 100 other plant species. By contrast, several guava species have become rare due to
habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and at least one (Jamaican guava, '' P. dumetorum''), is already extinct. Guava wood is used for meat smoking in Hawaii, and is used at barbecue competitions across the United States. In Cuba and Mexico, the leaves are used in barbecues.


Fruit

Guava fruits, usually long, are round or oval depending on the species. They have a pronounced and typical fragrance, similar to lemon rind but less sharp. The outer skin may be rough, often with a bitter taste, or soft and sweet. Varying between species, the skin can be any thickness, is usually green before maturity, but may be yellow, maroon, or green when ripe. The pulp inside may be sweet or sour and off-white ("white" guavas) to deep pink ("red" guavas). The seeds in the central pulp vary in number and hardness, depending on species.


Production

In 2019, world production of guavas was 55 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton ( United State ...
s, led by
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
with 45% of the total (table). Other major producers were China and Thailand.


Culinary uses

In Mexico and other Latin American countries, the popular beverage
agua fresca ''Aguas frescas'' ( en, cool waters, lit=fresh waters, italic=yes) are light non-alcoholic beverages made from one or more fruits, cereals, flowers, or seeds blended with sugar and water. They are popular in Mexico and some other Latin Ame ...
is often made with guava. The entire fruit is a key ingredient in punch, and the juice is often used in culinary sauces (hot or cold), ales, candies, dried snacks, fruit bars, and desserts, or dipped in chamoy. ''
Pulque Pulque (; nci, metoctli), or octli, is an alcoholic beverage made from the fermented sap of the maguey (agave) plant. It is traditional in central Mexico, where it has been produced for millennia. It has the color of milk, a rather viscous co ...
de guayaba'' ("guayaba" is Spanish for guava) is a popular alcoholic beverage in these regions. In many countries, guava is eaten raw, typically cut into quarters or eaten like an apple; it is also eaten with a pinch of salt and pepper, cayenne powder or a mix of spices (masala). In the Philippines, ripe guava is used in cooking ''
sinigang ''Sinigang'' is a Filipino soup or stew characterized by its sour and savory taste. It is most often associated with tamarind (Filipino: ''sampalok''), although it can use other sour fruits and leaves as the souring agent. It is one of the more ...
''. Guava is a popular snack in Cuba as ''pastelitos de guayaba''; and in Taiwan, sold on many street corners and night markets during hot weather, accompanied by packets of dried plum powder mixed with sugar and salt for dipping. In east Asia, guava is commonly eaten with sweet and sour dried plum powder mixtures. Guava juice is popular in many countries. The fruit is also often included in fruit salads. Because of its high level of
pectin Pectin ( grc, πηκτικός ': "congealed" and "curdled") is a heteropolysaccharide, a structural acid contained in the primary lamella, in the middle lamella, and in the cell walls of terrestrial plants. The principal, chemical component of ...
, guavas are extensively used to make candies,
preserves Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the method ...
, jellies,
jams Jams or JAMS may refer to: *Plural form of jam, a type of fruit preserve *Jams (clothing line) *JAMS (organization), United States organization that provides alternative dispute resolution services *The JAMs, former name of The KLF, a British band ...
, and marmalades (such as Brazilian ''
goiabada Goiabada (; from ''goiaba'', guava] is a Jam, conserve made of red guavas and sugar, commonly found throughout the Portuguese-speaking countries of the world. It dates back to the colonial times of Brazil, where guavas were used as a substitute ...
'' and Colombian and Venezuelan '' bocadillo (dessert), bocadillo''), and as a marmalade jam served on toast. Red guavas can be used as the base of salted products such as sauces, substituting for
tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
es, especially to minimize the acidity. A drink may be made from an
infusion Infusion is the process of extracting chemical compounds or flavors from plant material in a solvent such as water, oil or alcohol, by allowing the material to remain suspended in the solvent over time (a process often called steeping). An inf ...
of guava fruits and leaves, which in Brazil is called ''chá-de-goiabeira'', i.e., "tea" of guava tree leaves, considered medicinal.


Constituents


Nutrients

Guavas are rich in
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
and
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) an ...
, with moderate levels of
folic acid Folate, also known as vitamin B9 and folacin, is one of the B vitamins. Manufactured folic acid, which is converted into folate by the body, is used as a dietary supplement and in food fortification as it is more stable during processing and ...
(nutrition table). Low in
food energy Food energy is chemical energy that animals (including humans) derive from their food to sustain their metabolism, including their muscle, muscular activity. Most animals derive most of their energy from aerobic respiration, namely combining the ...
per typical serving, and with few
essential nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
s, a single common guava ('' P. guajava'') fruit contains 257% of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of health ...
(DV) for vitamin C (table). Nutrient content varies across guava
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s. Although the
strawberry guava ''Psidium cattleyanum (World Plants : Psidium cattleianum)'', commonly known as Cattley guava, strawberry guava or cherry guava, is a small tree (2–6 m tall) in the Myrtaceae (myrtle) family. The species is named in honour of English hor ...
(''P. littorale'' var. ''cattleianum'') has only 39% of the vitamin C in common varieties, its content in a 100 gram serving (90 mg) still provides 100% of the DV.


Phytochemicals

Guava leaves contain both
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
s and
polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some of ...
s like (+)-gallocatechin and
leucocyanidin Leucocyanidin is a colorless chemical compound that is a member of the class of natural products known as leucoanthocyanidins. Chemistry (+)-Leucocyanidin can be synthesized from (+)-dihydroquercetin by reduction with sodium borohydride. Mola ...
. As some of these
phytochemicals Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poison ...
produce the fruit skin and flesh color, guavas that are red-orange tend to have more polyphenol and carotenoid content than yellow-green ones.


Guava seed oil

Guava seed oil, which may be used for
culinary Culinary arts are the cuisine arts of outline of food preparation, food preparation, cooking and food presentation, presentation of food, usually in the form of meals. People working in this field – especially in establishments such as res ...
or
cosmetics Cosmetics are constituted mixtures of chemical compounds derived from either natural sources, or synthetically created ones. Cosmetics have various purposes. Those designed for personal care and skin care can be used to cleanse or protect ...
products, is a source of
beta carotene Beta (, ; uppercase , lowercase , or cursive Greek, cursive ; grc, βῆτα, bē̂ta or ell, βήτα, víta) is the second letter of the Greek alphabet. In the system of Greek numerals, it has a value of 2. In Modern Greek, it represents th ...
,
vitamin A Vitamin A is a fat-soluble vitamin and an essential nutrient for humans. It is a group of organic compounds that includes retinol, retinal (also known as retinaldehyde), retinoic acid, and several provitamin A carotenoids (most notably bet ...
,
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) an ...
,
copper Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...
,
zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...
and
selenium Selenium is a chemical element with the symbol Se and atomic number 34. It is a nonmetal (more rarely considered a metalloid) with properties that are intermediate between the elements above and below in the periodic table, sulfur and tellurium, ...
, and is particularly rich in
linoleic acid Linoleic acid (LA) is an organic compound with the formula COOH(CH2)7CH=CHCH2CH=CH(CH2)4CH3. Both alkene groups are cis-trans isomerism, ''cis''. It is a fatty acid sometimes denoted 18:2 (n-6) or 18:2 ''cis''-9,12. A linoleate is a salt (chem ...
. The composition of fatty acids in guava seed oil is presented in the following table:


Folk medicine

Since the 1950s, guavas – particularly the leaves – have been studied for their constituents, potential biological properties and history in
folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
.


Parasites

Guavas are one of the most common hosts for fruit flies like '' A. suspensa'', which lay their eggs in overripe or spoiled guavas. The larvae of these flies then consume the fruit until they can proceed into the
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
stage. This
parasitism Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
has led to millions in economic losses for nations in Central America.


See also

* Myrteae, the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
containing guava and closely related plants with fleshy fruit


References

{{Reflist Berries Edible fruits Mexican cuisine Cuban cuisine Belizean cuisine