''Gryponyx'' (meaning "hooked-claw") is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

massopod Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defi ...

sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...

known from southern Free State, central

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

.Broom, R. (1911). On the dinosaurs of the Stormberg, South Africa. ''Annals of the South African Museum'' 7(4):291-308.

Description

''Gryponyx africanus'' is known from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

SAM 3357-59, a nearly complete

postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

which includes partial

vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordata, ...

pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

, both

forelimb A forelimb or front limb is one of the paired articulated appendages (limbs) attached on the cranial ( anterior) end of a terrestrial tetrapod vertebrate's torso. With reference to quadrupeds, the term foreleg or front leg is often used instead. ...

s and both

hindlimb A hindlimb or back limb is one of the paired articulated appendages (limbs) attached on the caudal ( posterior) end of a terrestrial tetrapod vertebrate's torso.http://www.merriam-webster.com/medical/hind%20limb, Merriam Webster Dictionary-Hindl ...

s. ''Gryponyx'' has been estimated to have been about in length. It was collected from the

Upper Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, sout ...

of the

Stormberg Group The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group roc ...

(

Karoo Basin The Karoo Supergroup is the most widespread stratigraphy, stratigraphic unit in Africa south of the Kalahari Desert. The group (stratigraphy), supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Pennsylvan ...

), dating to the

Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...

Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian and is ...

stages of the

Lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma ...

period. It was originally described by Broom (1911) as a

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

. Huene (1932) named the family Gryponychidae to contain ''Gryponyx'' and ''Aetonyx'' and placed it within

Carnosauria Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Gig ...

.Friedrich von Huene (1932) Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte. ''Monographien zur Geologie und Paläontologie, Series 1'' 4: 1–361

Galton Sir Francis Galton, FRS FRAI (; 16 February 1822 – 17 January 1911), was an English Victorian era polymath: a statistician, sociologist, psychologist, anthropologist, tropical explorer, geographer, inventor, meteorologist, proto-gene ...

and Cluver synonymized ''G. africanus'' with '' Massospondylus harriesi'' in 1976,Galton, P.M., and Cluver, M.A. (1976). ''Anchisaurus capensis'' (Broom) and a revision of the Anchisauridae (Reptilia, Saurischia). ''Annals of the South African Museum'' 69(6):121-159. which was in turn synonymized by

Michael Cooper Michael Jerome Cooper (born April 15, 1956) is an American basketball coach and former player who is the boys varsity coach at Culver City High School. He played for the Los Angeles Lakers in the National Basketball Association (NBA), winning ...

in 1981 with ''

Massospondylus carinatus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by Si ...

''Cooper, M.R. (1981). The prosauropod dinosaur ''Massospondylus carinatus'' Owen from Zimbabwe: its biology, mode of life and phylogenetic significance. ''Occasional Papers of the National Museums and Monuments of Rhodesia, Series B, Natural Sciences'' 6(10):689-840. (and today ''M. harriesi'' is considered to be a ''

nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...

'').Galton, P.M., and Upchurch, P. (2004). Prosauropoda. In: D. B. Weishampel, P. Dodson, & H. Osmólska (eds.), ''The Dinosauria'' (second edition). University of California Press:Berkeley, 232-258. However, Vasconcelos and Yates (2004) found ''Gryponyx'' to be distinctive enough from other basal sauropodomorphs to be placed in its own genus. They found that it differs from other taxa by the following characteristics: total length of metacarpal I exceeds maximum proximal width and a long, narrow pubic apron with straight lateral margins. Although this publication wasn't formal, they conducted a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis using Yates (2004) sauropodomorph matrix and found ''Gryponyx'' to be the most basal massospondylid.C. C. Vasconcelos, A. M. Yates (2004). Sauropodomorph biodiversity of the upper Elliot Formation (Lower Jurassic) of southern Africa. ''Geoscience Africa, Abstract Volume'' 2:670. The same result was found by

Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...

''et al.'' (2010). Yates ''et al.'' (2010) recovered ''Gryponyx'' in a trichotomy with

Massospondylidae Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarctica." Geologica ...

and Anchisauria. However, ''Gryponyx'' has yet to be formally redescribed. Two additional species of ''Gryponyx'' have been described: ''G. transvaalensis'' was described on the basis of finger bones and the anterior limb metatarsals from the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...

Bushveld Sandstone Formation The Bushveld Sandstone is a geological formation dating to roughly between 201 and 189 million years ago and covering the Carnian to Norian stages. The Bushveld Sandstone is found in Transvaal, South Africa and is a member of the Stormberg Group. ...

Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name Transvaal. * South African Republic (1856–1902; af, ...

. ''G. taylori'' was described on the basis of

sacral Sacral may refer to: *Sacred, associated with divinity and considered worthy of spiritual respect or devotion *Of the sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spi ...

and

pelvic The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

rim from the Upper Elliot Formation, southern Free State. Galton and Cluver (1976) synonymized ''G. taylori'' with ''M. harriesi'' and considered ''G. transvaalensis'' to be a ''nomen dubium''. Both ''G. taylori'' and ''G. transvaalensis'' were synonymized by Michael Cooper (1981) with ''M. carinatus'' and Galton and Upchurch (2004) considered them to be dubious.

Etymology

''Gryponyx'' was first named by

Robert Broom Robert Broom FRS FRSE (30 November 1866 6 April 1951) was a British- South African doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University of Glasgow. From 1903 to 1910, he ...

1911 A notable ongoing event was the Comparison of the Amundsen and Scott Expeditions, race for the South Pole. Events January * January 1 – A decade after federation, the Northern Territory and the Australian Capital Territory ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Gryponyx africanus''. The generic name is derived from ''grypos'',

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for "hooked" and ''onyx'', Greek for "claw". The specific name refers to

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, where the holotype was discovered.

References

{{Taxonbar, from=Q1989760 Sauropodomorphs Hettangian life Pliensbachian life Sinemurian life Early Jurassic dinosaurs of Africa Jurassic South Africa Fossils of South Africa Fossil taxa described in 1911 Taxa named by Robert Broom