Grimmia Rigidissima
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''Racomitrium lanuginosum'' is a widespread species of
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
found in montane and arctic
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
, the genus Racomitrium is found across the Northern and Southern hemispheres., however Racomitrium lanuginosum is only found in the Northern hemisphere. It grows as large mats on exposed rock and in boulder scree, particularly on acidic rocks. Its
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
have a characteristically
decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...
and toothed hair-point, which gives rise to its regional common names woolly fringemoss, hoary rock-moss and woolly moss.


Description

''Racomitrium lanuginosum'' grows as "large cushiony mats". Its stems are up to long and irregularly branched. The leaves are long, wide,
lanceolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
, pointed and with a strong
midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
. They end in a long, thin hair-point, with teeth along both sides pointing 40°–90° from the axis of the leaf. The hyaline (transparent) margins of the hair-point are
decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...
along the sides of the leaf. Especially when dry, these hair-points give the plant a downy appearance, which is referenced in several of the common names for the species.
Sporophyte A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote pr ...
s are rarely produced, although they are reported to be abundant in coastal areas. The capsules are long, smooth and brown, and contain spores, which are 8–12 Î¼m long. The form of the hair-points on the leaves of ''R. lanuginosum'' is unique among mosses.


Similar species

''Racomitrium lanuginosum'' is "one of the few bryophytes familiar to many non-bryologists". Other species that may be confused with it include '' R. canescens'' and its close relatives, '' R. heterostichum'' and ''
Hedwigia stellata ''Hedwigia'' is a genus of mosses belonging to the family Hedwigiaceae. The genus was first described by Palisot de Beauvois in 1804. The genus has cosmopolitan distribution. Species: * ''Hedwigia ciliata ''Hedwigia ciliata'' is a species ...
''.


Distribution and ecology

''Racomitrium lanuginosum'' grows from
sea level Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardised g ...
to the
alpine zone Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alpine tundra gets ...
, particularly among boulder scree and on exposed rock surfaces. It will also grow in
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
or
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...
s. It is more frequently found on acidic rocks than calcareous ones. ''Racomitrium lanuginosum'' has a
circumpolar distribution A circumpolar distribution is any range of a taxon that occurs over a wide range of longitudes but only at high latitudes; such a range therefore extends all the way around either the North Pole or the South Pole. Taxa that are also found in isolat ...
in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, and is found disjunctly in the mountains of the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and the Southern Hemisphere. Within North America, ''R. lanuginosum'' is found widely across northern Canada, and extends further south both in the east (to
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
), and in the west (to
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
and the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
). It is also found in
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
and in mountainous regions of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
and
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. In the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
, isolated populations occur on
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
, the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
,
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
,
Tristan da Cunha Tristan da Cunha (), colloquially Tristan, is a remote group of volcanic islands in the South Atlantic Ocean. It is the most remote inhabited archipelago in the world, lying approximately from Cape Town in South Africa, from Saint Helena ...
and
South Georgia South Georgia ( es, Isla San Pedro) is an island in the South Atlantic Ocean that is part of the British Overseas Territory of South Georgia and the South Sandwich Islands. It lies around east of the Falkland Islands. Stretching in the east†...
; it also occurs on
Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...
in the Indian Ocean, in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
and the sub-Antarctic islands
Deception Island Deception Island is an island in the South Shetland Islands close to the Antarctic Peninsula with a large and usually "safe" natural harbor, which is occasionally troubled by the underlying active volcano. This island is the caldera of an act ...
,
Kerguelen The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a archipelago, group of islands in the subantarctic, sub-Antarctic constituting one of the two exposed parts of the ...
,
ÃŽles Crozet The Crozet Islands (french: ÃŽles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic ...
,
Heard Island The Territory of Heard Island and McDonald Islands (HIMI) is an Australian external territory comprising a volcanic group of mostly barren Antarctic islands, about two-thirds of the way from Madagascar to Antarctica. The group's overall size ...
and the
Prince Edward Islands The Prince Edward Islands are two small uninhabited islands in the sub-Antarctic Indian Ocean that are part of South Africa. The islands are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and Prince Edward Island ...
. As well as occurring widely across Arctic Asia, ''R. lanuginosum'' is also found in some temperate and tropical mountains on
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
,
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
. In Australia, ''R. lanuginosum'' is restricted to
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
and the highest ground of the
Great Dividing Range The Great Dividing Range, also known as the East Australian Cordillera or the Eastern Highlands, is a cordillera system in eastern Australia consisting of an expansive collection of mountain ranges, plateaus and rolling hills, that runs rough ...
on the border of
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
and
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
.


Taxonomy

''Racomitrium lanuginosum'' was first described by
Johann Hedwig Johann Hedwig (8 December 1730 – 18 February 1799), also styled as Johannes Hedwig, was a German botanist notable for his studies of mosses. He is sometimes called the "father of bryology". He is known for his particular observations of sexual r ...
in 1801 as ''Trichostomum lanuginosum''. It was moved to its current genus in 1818 when
Samuel Elis̩e Bridel-Brideri Samuel Elis̩e Bridel-Brideri (28 November 1761 in Crassier, Vaud Р7 January 1828) was a Swiss-German bryologist. He studied at the University of Lausanne, and at the age of 19 began work as a tutor to the princes of Saxe-Gotha-Altenburg. ...
established the genus ''Racomitrium'', and that position was cemented in 1860 when
Wilhelm Philippe Schimper Wilhelm Philippe Schimper (January 12, 1808 – March 20, 1880, in Lichtenberg) was an Alsatian botanist with French, later German citizenship. He was born in Dossenheim-sur-Zinsel, but spent his youth in Offwiller, a village at the foot of the ...
designated ''R. lanuginosum'' as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of ''Racomitrium''. ''Racomitrium lanuginosum'' is classified in ''Racomitrium'' sect. ''Racomitrium'', a group that also contains the species '' R. geronticum'' and '' R. pruinosum''. ''Racomitrium lanuginosum'' is known by a variety of
vernacular name A vernacular or vernacular language is in contrast with a "standard language". It refers to the language or dialect that is spoken by people that are inhabiting a particular country or region. The vernacular is typically the native language, n ...
s around the world. In the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
, it is known as the "woolly fringe-moss", in Canada as the "hoary rock-moss", and in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
as "woolly moss".


References


External links

* {{Taxonbar, from=Q3024247 Grimmiales Plants described in 1801