Grielum Grandiflorum
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''Grielum'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of four accepted species of plants in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Neuradaceae The Neuradaceae are a family of flowering plant, comprising three genera — ''Grielum'', '' Neurada'' and '' Neuradopsis'' — totalling ten known species. These genera were formerly placed in order Rosales, and in one case even in family Rosa ...
. They are low-growing annual herbs endemic to dry regions, mainly in Western
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
and
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
.


Description

The genus comprises annual dwarf herbs with pronounced
taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...
s. * The leaves are more or less furry, and may be canescent or
tomentose Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...
. They are
lobed The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
at least, but more typically are decompound. The stipules are
obsolete Obsolescence is the state of being which occurs when an object, service, or practice is no longer maintained or required even though it may still be in good working order. It usually happens when something that is more efficient or less risky r ...
, being barely evident if present at all. * The flowers are conspicuous and are borne on singly on
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
s in leaf axils. They are generally vivid lemon-yellow in colour, but with a white centre in some species. The calyx has five sepals, forming a shallow cup. The corolla has five petals, usually imbricate and thereby forming a shallow bowl, though this is less evident if the petals happen to be narrow, as sometimes occurs on a particular plant. The petals are inserted into the throat of the calyx cup. There are ten stamens, more or less set in two whorls of five, inserted between the base of the gynoecium and the petals. The carpels typically are five in number, but may vary from about 3 to 10. They are connate and remain so in the fruit, which forms a synaptospermic
diaspore Diaspore , also known as diasporite, empholite, kayserite, or tanatarite, is an aluminium oxide hydroxide mineral, α-AlO(OH), crystallizing in the orthorhombic system and isomorphous with goethite. It occurs sometimes as flattened crystals, bu ...
that may act as a trample burr, being spread by sticking to animals' feet. The
style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...
s are
terete Terete is a term in botany used to describe a cross section that is circular, or like a distorted circle, with a single surface wrapping around it.Lichen Vocabulary, Lichens of North America Information, Sylvia and Stephen Sharnoff/ref> This is us ...
and the stigmas
capitate The capitate bone is a bone in the human wrist found in the center of the carpal bone region, located at the distal end of the radius and ulna bones. It articulates with the third metacarpal bone (the middle finger) and forms the third carpomet ...
. * The taproot of local species is well developed as a storage organ and is an important source of food and moisture for some animals such as
duiker A duiker is a small to medium-sized brown antelope native to sub-Saharan Africa, found in heavily wooded areas. The 22 extant species, including three sometimes considered to be subspecies of the other species, form the subfamily Cephalophina ...
, so much so that one of its common names is "''duikerwortel''",
Afrikaans Afrikaans (, ) is a West Germanic language that evolved in the Dutch Cape Colony from the Dutch vernacular of Holland proper (i.e., the Hollandic dialect) used by Dutch, French, and German settlers and their enslaved people. Afrikaans gra ...
for "duiker root"; duikers are notorious for digging up the root from sandy soils. It also was an important food plant for local hunter-gatherers. It is thick and so mucilaginous that it also is known by variations on the name "Pietsnot", meaning "snotty Pete". The taproot of for example ''Grielum humifusum'' and ''Grielum grandiflorum'' are relished even by some campers when roasted on a campfire.Roasted roots
/ref>


Species

Four species are recognised: * ''
Grielum cuneifolium ''Grielum'' is a genus of four accepted species of plants in the family Neuradaceae. They are low-growing annual herbs endemic to dry regions, mainly in Western South Africa and Namibia. Description The genus comprises annual dwarf herbs with ...
'' Schinz * ''
Grielum grandiflorum ''Grielum'' is a genus of four accepted species of plants in the family Neuradaceae. They are low-growing annual herbs endemic to dry regions, mainly in Western South Africa and Namibia. Description The genus comprises annual dwarf herbs with ...
'' (L.) Druce * ''
Grielum humifusum ''Grielum humifusum'' is a species of flowering plant in the family Neuradaceae The Neuradaceae are a family of flowering plant, comprising three genera — ''Grielum'', '' Neurada'' and '' Neuradopsis'' — totalling ten known species. Thes ...
'' Thunb. * ''
Grielum sinuatum ''Grielum'' is a genus of four accepted species of plants in the Family (biology), family Neuradaceae. They are low-growing annual herbs endemic to dry regions, mainly in Western South Africa and Namibia. Description The genus comprises annual ...
'' Licht. ex Burch.


References

Neuradaceae Malvales genera {{Malvales-stub