Greater Shearwater
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The great shearwater (''Ardenna gravis'') is a large
shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwa ...
in the
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
family
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which also ...
. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic.


Taxonomy

The great shearwater was formally described in 1818 by the Irish naturalist Bernard O'Reilly and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Procellaria gravis''. The great shearwater is now placed in the genus ''
Ardenna ''Ardenna'' is a genus of seabirds in the family Procellariidae. These medium-sized shearwater species were formerly included in the genus ''Puffinus''. Taxonomy A phylogenetic analysis using mitochondrial DNA published in 2004 found that ''Puff ...
'' that was introduced in 1853 by
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
. The genus name ''Ardenna'' was used to refer to a seabird by Italian naturalist
Ulisse Aldrovandi Ulisse Aldrovandi (11 September 1522 – 4 May 1605) was an Italian naturalist, the moving force behind Bologna's botanical garden, one of the first in Europe. Carl Linnaeus and the comte de Buffon reckoned him the father of natural history st ...
in 1603. The specific epithet ''gravis'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
meaning "heavy" or "weighty". The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
: no subspecies are recognised.


Description

This shearwater is 43–51 cm (16.9–20.1 in) in length with a 105–122 cm (3.5–4.0 ft) wingspan. It is identifiable by its size, dark upper parts, and white under parts, with the exception of a brown belly patch and dark shoulder markings. It has a black cap, black bill, and a white "horseshoe" on the base of the tail. The stiff flight, like a large
Manx shearwater The Manx shearwater (''Puffinus puffinus'') is a medium-sized shearwater in the seabird family Procellariidae. The scientific name of this species records a name shift: Manx shearwaters were called Manks puffins in the 17th century. Puffin is an ...
, is also distinctive. The only other large shearwaters in its range are the
Cory's Shearwater Cory's shearwater (''Calonectris borealis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially of rocky islands in the eastern Atlantic. Outside the breeding season it ranges widely in the Atlantic. It was formerly ...
and the all-dark
Sooty Shearwater The sooty shearwater (''Ardenna grisea'') is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name , and as muttonbird, like its relatives the wedge-tailed shearwater (''A. pacificus' ...
. Greater shearwater (Ardenna gravis) in flight Sagres 2.jpg, wing upperside Greater shearwater (Ardenna gravis) in flight Sagres.jpg, wing underside


Distribution and habitat

The great shearwater, like the
sooty shearwater The sooty shearwater (''Ardenna grisea'') is a medium-large shearwater in the seabird family Procellariidae. In New Zealand, it is also known by its Māori name , and as muttonbird, like its relatives the wedge-tailed shearwater (''A. pacificus' ...
, follows a circular migration route, moving north up the eastern seaboard of first
South South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and then
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, before crossing the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
in August. It can be quite common off the southwestern coasts of Great Britain and Ireland before heading back south again, this time down the eastern
littoral The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas ...
of the Atlantic.


Behaviour and ecology

This bird has the typically "shearing" flight of the genus, dipping from side to side on stiff wings with few wingbeats, the wingtips almost touching the water. Its flight is powerful and direct, with wings held stiff and straight.


Breeding

This species breeds on
Nightingale Island Nightingale Island is an active volcanic island in the South Atlantic Ocean, in area, part of the Tristan da Cunha group of islands. They are administered by the United Kingdom as part of the British overseas territory, overseas territory of Sai ...
,
Inaccessible Island Inaccessible Island is a volcanic island located in the South Atlantic Ocean, south-west of Tristan da Cunha. Its highest point, Cairn Peak, reaches , and the island is in area. The volcano was last active six million years ago and is curren ...
,
Tristan da Cunha Tristan da Cunha (), colloquially Tristan, is a remote group of volcanic islands in the South Atlantic Ocean. It is the most remote inhabited archipelago in the world, lying approximately from Cape Town in South Africa, from Saint Helena ...
, and
Gough Island upright=1.3, Map of Gough island Gough Island ( ), also known historically as Gonçalo Álvares, is a rugged volcanic island in the South Atlantic Ocean. It is a dependency of Tristan da Cunha and part of the British overseas territory of Sain ...
. It is one of only a few
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
species to
migrate Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
from breeding grounds in the Southern Hemisphere to the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, the normal pattern being the other way around. This shearwater nests in large colonies, laying one white egg in a small burrow or in the open grass. These nests are visited only at night to avoid predation by large
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...
s.


Food and feeding

The great shearwater feeds on
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
and
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
, which it catches from the surface or by plunge-diving. It readily follows fishing boats, where it indulges in noisy squabbles. This is a gregarious species, which can be seen in large numbers from ships or appropriate headlands. They have a piercing ''"eeyah''" cry usually given when resting in groups on the water. Great shearwaters are among the seabird species with the highest incidence of plastic ingestion.


References


Further reading

*Austin, Jeremy J.; Bretagnolle, Vincent & Pasquet, Eric (2004): A global molecular phylogeny of the small ''Puffinus'' shearwaters and implications for systematics of the Little-Audubon's Shearwater complex. ''
Auk An auk or alcid is a bird of the family Alcidae in the order Charadriiformes. The alcid family includes the murres, guillemots, auklets, puffins, and murrelets. The word "auk" is derived from Icelandic ''álka'', from Old Norse ''alka'' (a ...
'' 121(3): 847–864. DOI: 10.1642/0004-8038(2004)121 847:AGMPOT.0.CO;2HTML abstract
*Bull, John L.; Farrand, John Jr.; Rayfield, Susan &
National Audubon Society The National Audubon Society (Audubon; ) is an American non-profit environmental organization dedicated to conservation of birds and their habitats. Located in the United States and incorporated in 1905, Audubon is one of the oldest of such organ ...
(1977): ''The Audubon Society field guide to North American birds, Eastern Region''. Alfred A. Knopf, New York. *Harrison, Peter (1987): ''Seabirds of the world : a photographic guide''. Princeton University Press, Princeton.


External links


Greater shearwater photos


{{Taxonbar, from=Q27074609
great shearwater The great shearwater (''Ardenna gravis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic. Taxonomy The great s ...
Birds of the Atlantic Ocean Birds of islands of the Atlantic Ocean Birds of subantarctic islands Fauna of Gough Island
great shearwater The great shearwater (''Ardenna gravis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic. Taxonomy The great s ...
great shearwater The great shearwater (''Ardenna gravis'') is a large shearwater in the seabird family Procellariidae. It breeds colonially on rocky islands in the south Atlantic. Outside the breeding season it ranges widely in the Atlantic. Taxonomy The great s ...