Greater Sandhill Crane
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The sandhill crane (''Antigone canadensis'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of large crane of North America and extreme northeastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. The common name of this bird refers to
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
like that at the
Platte River The Platte River () is a major river in the State of Nebraska. It is about long; measured to its farthest source via its tributary, the North Platte River, it flows for over . The Platte River is a tributary of the Missouri River, which itself ...
, on the edge of
Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the southwe ...
's Sandhills on the American
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, an ...
. Sandhill Cranes are known to hangout at the edges of bodies of water especially in the Central Florida region. This is the most important stopover area for the nominotypical
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, the lesser sandhill crane (''A. c. canadensis''), with up to 450,000 of these birds migrating through annually.


Taxonomy

In 1750, English naturalist George Edwards included an illustration and a description of the sandhill crane in the third volume of his ''A Natural History of Uncommon Birds''. He used the English name "The Brown and Ash-colour'd Crane". Edwards based his hand-coloured etching on a preserved specimen that had been brought to London from the
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
area of Canada by
James Isham James Isham (1716–1761) was chief factor (master) at both York Factory and Fort Prince of Wales in Canada during the mid-1700s. He kept detailed journals that described life in the region, including flora and fauna that were unknown to people in ...
. When in 1758 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the tenth edition, he placed the sandhill crane with herons and cranes in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Ardea''. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Ardea canadensis'', and cited Edwards' work. The sandhill crane was formerly placed in the genus '' Grus'', but a
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2010 found that the genus, as then defined, was
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
. In the resulting rearrangement to create
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
genera, four species, including the sandhill crane, were placed in the resurrected genus ''
Antigone In Greek mythology, Antigone ( ; Ancient Greek: Ἀντιγόνη) is the daughter of Oedipus and either his mother Jocasta or, in another variation of the myth, Euryganeia. She is a sister of Polynices, Eteocles, and Ismene.Roman, L., & Roma ...
'' that had originally been erected by the German naturalist
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
in 1853. The specific epithet ''canadensis'' is the
modern Latin New Latin (also called Neo-Latin or Modern Latin) is the revival of Literary Latin used in original, scholarly, and scientific works since about 1500. Modern scholarly and technical nomenclature, such as in zoological and botanical taxonomy ...
word for "from Canada". Five
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''A. c. canadensis'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758) – northeast Siberia through Alaska and north Canada to
Baffin Island Baffin Island (formerly Baffin Land), in the Canadian territory of Nunavut, is the largest island in Canada and the fifth-largest island in the world. Its area is , slightly larger than Spain; its population was 13,039 as of the 2021 Canadia ...
* ''A. c. tabida'' ( Peters, JL, 1925) – south Canada and west, central United States * ''A. c. pratensis'' ( Meyer, FAA, 1794) – Georgia and Florida * ''A. c. pulla'' (
Aldrich Aldrich may refer to: Places United States *Aldrich, Alabama, unincorporated community *Aldrich, Minnesota, city *Aldrich Township, Wadena County, Minnesota *Aldrich, Missouri, village People *Aldrich (surname), a surname (including a list of pe ...
, 1972) – Mississippi * ''A. c. nesiotes'' (
Bangs Bang or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Lorestan, a village in I ...
& Zappey, 1905) – Cuba and
Isla de la Juventud Isla de la Juventud (; en, Isle of Youth) is the second-largest Cuban island (after Cuba's mainland) and the seventh-largest island in the West Indies (after mainland Cuba itself, Hispaniola, Jamaica, Puerto Rico, Trinidad, and Andros Islan ...
(Isle of Pines)


Description

Adults are gray overall; during breeding, their
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is usually much worn and stained, particularly in the migratory populations, and looks nearly
ochre Ochre ( ; , ), or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the colours produced ...
. The average weight of the larger males is , while the average weight of females is , with a range of across the subspecies. Sandhill cranes have red foreheads, white cheeks, and long, dark, pointed
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
s. In flight, their long, dark legs trail behind, and their long necks keep straight. Immature birds have reddish-brown upperparts and gray underparts. The sexes look alike. Sizes vary among the different
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
; the average height of these birds is around .Dunn, J.L. & Alderfer, J., editors. Field Guide to the Birds of North America. National Geographic, Washington, D.C. Their wing chords are typically , tails are , the exposed culmens are long, and the tarsi measure . Wingspan is 78.7 in (200 cm). These cranes frequently give a loud, trumpeting call that suggests a rolled "r" in the throat, and they can be heard from a long distance. Mated pairs of cranes engage in "unison calling". The cranes stand close together, calling in a synchronized and complex duet. The female makes two calls for every one from the male. Sandhill cranes' large wingspans, typically , make them very skilled
soaring Soaring may refer to: * Gliding, in which pilots fly unpowered aircraft known as gliders or sailplanes * Lift (soaring), a meteorological phenomenon used as an energy source by some aircraft and birds * ''Soaring'' (magazine), a magazine produced ...
birds, similar in style to
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s and
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s. Using
thermal A thermal column (or thermal) is a rising mass of buoyant air, a convective current in the atmosphere, that transfers heat energy vertically. Thermals are created by the uneven heating of Earth's surface from solar radiation, and are an example ...
s to obtain lift, they can stay aloft for many hours, requiring only occasional flapping of their wings, thus expending little energy. Migratory flocks contain hundreds of birds, and can create clear outlines of the normally invisible rising columns of air (thermals) they ride. Sandhill cranes fly south for the winter. In their wintering areas, they form flocks over 10,000. One place this happens is at
Bosque del Apache National Wildlife Refuge The Bosque del Apache National Wildlife Refuge ( ) is located in southern New Mexico. It was founded in 1939 and is administered by the U.S. Fish and Wildlife Service. It is a favorite spot to watch the migration of the sandhill cranes in th ...
, south of
Albuquerque, New Mexico Albuquerque ( ; ), ; kee, Arawageeki; tow, Vakêêke; zun, Alo:ke:k'ya; apj, Gołgéeki'yé. abbreviated ABQ, is the most populous city in the U.S. state of New Mexico. Its nicknames, The Duke City and Burque, both reference its founding in ...
. An annual Sandhill Crane Festival is held there in November. File:Grus canadensis in flight-2618.jpg, Sandhill crane in flight at the Llano Seco Unit of the
Sacramento National Wildlife Refuge Complex The Sacramento National Wildlife Refuge Complex is part of the United States system of National Wildlife Refuges (NWR). It is located in northern California, in the valley of the Sacramento River. The Sacramento NWR Complex was created in an att ...
, California, USA File:Immature Florida Sandhill.jpg, An immature Florida sandhill crane walks along the shore of Lake Cecile near
Kissimmee, Florida Kissimmee ( ) is the largest city and county seat of Osceola County, Florida, United States. As of the 2020 census, the population was 79,226. It is a Principal City of the Orlando-Kissimmee-Sanford, Florida, Metropolitan Statistical Area, wh ...
. File:Sandhill crane landing at Llano Seco-9853.jpg, Sandhill crane in flight at the Sacramento National Wildlife Refuge File:Two Florida Sandhill Cranes at a gas station near Cape Canaveral, Florida.jpg, Two Florida sandhill cranes at a gas station near Cape Canaveral, Florida


Fossil record

Sandhill cranes have one of the longest
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
histories of any extant bird. A 10-million-year-old crane fossil from
Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the southwe ...
is said to be of this
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, but this may be from a prehistoric relative or ancestor of sandhill cranes, and not belong in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Grus''. The oldest unequivocal sandhill crane fossil is 2.5 million years old, older by half than the earliest remains of most living species of birds, primarily found from after the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
boundary some 1.8 million years ago. As these ancient sandhill cranes varied as much in size as present-day birds, those Pliocene fossils are sometimes described as new species. ''Grus haydeni'' may have been a prehistoric relative, or it may comprise material of a sandhill crane and its ancestor.


Subspecies and evolution

Sandhill cranes vary considerably in size (much of which is clinal) and in migratory habits. A female of ''A. c. canadensis'' averages , in length, and has a wingspan of . A male of ''A. c. tabida'' averages , in length, and has a wingspan of . The southern subspecies (along with ''A. c. rowani'') are intermediate, roughly according to
Bergmann's rule Bergmann's rule is an ecogeographical rule that states that within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer ...
. Three subspecies are resident: ''A. c. pulla'' of the Gulf Coast of the U.S., ''A. c. pratensis'' of
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
and
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, and ''A. c. nesiotes'' of
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
. The northern populations exist as fragmented remains in the contiguous U.S. and a large and contiguous population from Canada to
Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
. These migrate to the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
and
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. These cranes are rare vagrants to China,
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
, and Japan and very rare vagrants to
Western Europe Western Europe is the western region of Europe. The region's countries and territories vary depending on context. The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
. Six subspecies have been recognized in recent times: *Lesser sandhill crane, ''A. c. canadensis'' *Cuban sandhill crane, ''A. c. nesiotes'' –
ESA , owners = , headquarters = Paris, Île-de-France, France , coordinates = , spaceport = Guiana Space Centre , seal = File:ESA emblem seal.png , seal_size = 130px , image = Views in the Main Control Room (1205 ...
: endangered *Florida sandhill crane, ''A. c. pratensis'' *Mississippi sandhill crane, ''A. c. pulla'' – ESA: endangered *Canadian sandhill crane, ''A. c. rowani'' *Greater sandhill crane, ''A. c. tabida'' The Florida sandhill crane was listed as EC or easily confused to facilitate an attempted reintroduction of the
whooping crane The whooping crane (''Grus americana'') is the tallest North American bird, named for its whooping sound. It is an endangered crane species. Along with the sandhill crane (''Antigone canadensis''), it is one of only two crane species native to N ...
(''Grus americana'') into Florida. The attempt failed, but the listing remained. The current list of endangered subspecies includes only two birds, ''A. c. nesiotes'' and ''A. c. rowani'', with ''A. c. pratensis'' no longer listed. Sandhill cranes occur in pastures, open prairies and freshwater wetlands in peninsular Florida from the Everglades to the Okefenokee Swamp. Some authorities no longer recognize Canadian sandhill crane as a distinct subspecies, as insignificant genetic differentiation and minimal morphological differentiation exist between it and greater sandhill crane. The others can be somewhat more reliably distinguished in hand by measurements and plumage details, apart from the size differences already mentioned. Unequivocal identification often requires location information, which is often impossible in migrating birds. Analysis of control region
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
haplotype A haplotype ( haploid genotype) is a group of alleles in an organism that are inherited together from a single parent. Many organisms contain genetic material ( DNA) which is inherited from two parents. Normally these organisms have their DNA or ...
data shows two major lineages. The
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
and the
subarctic The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
migratory population includes the lesser sandhill cranes. The other lineages can be divided into a migratory and some indistinct clusters which can be matched to the resident subspecies. The lesser and greater sandhill cranes are quite distinct, their divergence dating to roughly 2.3–1.2
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
, some time during the
Late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
or
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
.
Glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betw ...
seemingly fragmented off a founder population of lesser sandhill cranes, because during each major
ice age An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gree ...
, its present breeding range was frozen year-round. Still, sandhill cranes are amply documented from fossil and
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
remains right to the modern era. Conceivably, they might be considered distinct species already, a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
''G. canadensis'' and the greater sandhill crane, ''G. pratensis'', which would include the other populations. The scant differences between southern Canadian and western U.S. populations appear to result from
genetic drift Genetic drift, also known as allelic drift or the Wright effect, is the change in the frequency of an existing gene variant (allele) in a population due to random chance. Genetic drift may cause gene variants to disappear completely and there ...
, due to the recent reduction in population and range fragmentation. Until the early 20th century, the southern migratory birds occupied a much larger and continuous range. Thus, the subspecies ''A. c. rowani'' may well be abandoned. The two southern U.S. resident populations are somewhat more distinct. The Cuban population has been comparatively little studied, but appears to have been established on the island for a long time. They and the migratory greater sandhill cranes proper may form a group of lineages that diverged much later from a range in the southern U.S. and maybe northern Mexico, where they were resident. The southern migratory population would then represent a later re-expansion, which (re-) evolved their migratory habits independent from the northernmost birds, the geographically separated populations expanding rapidly when more habitat was available as the last ice age ended.


Behavior

Sandhill cranes are fairly social birds that usually live in pairs or family groups through the year. During migration and winter, unrelated cranes come together to form "survival groups" that forage and roost together. Such groups often congregate at migration and winter sites, sometimes in the thousands. Sandhill cranes are mainly herbivorous, but eat various types of food, depending on availability. They often feed with their bills down to the ground as they root around for seeds and other foods, in shallow wetlands with vegetation or various upland habitats. Cranes readily eat cultivated foods such as
corn Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...
,
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
,
cottonseed Cottonseed is the seed of the cotton plant. Composition The mature seeds are brown ovoids weighing about a tenth of a gram. By weight, they are 60% cotyledon, 32% coat and 8% embryonic root and shoot. These are 20% protein, 20% oil and 3.5% sta ...
, and
sorghum ''Sorghum'' () is a genus of about 25 species of flowering plants in the grass family (Poaceae). Some of these species are grown as cereals for human consumption and some in pastures for animals. One species is grown for grain, while many othe ...
. Waste corn is useful to cranes preparing for migration, providing them with nutrients for the long journey. Among northern races of sandhill cranes, their diet is most varied, especially among breeding birds. They variously feed on
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
, small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s,
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
s,
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s, and
amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...
s. Sandhill cranes raise one brood per year. In nonmigratory populations, laying begins between December and August. In migratory populations, laying usually begins in April or May. Both members of a breeding pair build the nest using plant material from the surrounding area. Nest sites are usually marshes, bogs, or swales, though occasionally on dry land. Females lay one to three (usually two) oval, dull brown eggs with reddish markings. Both parents incubate the eggs for about 30 days. The chicks are precocial; they hatch covered in down, with their eyes open, and able to leave the nest within a day. The parents brood the chicks for up to three weeks after hatching, feeding them intensively for the first few weeks, then gradually less frequently until they reach independence at 9 to 10 months old. The chicks remain with their parents until one to two months before the parents lay the next clutch of eggs the following year, remaining with them 10–12 months. After leaving their parents, the chicks form nomadic flocks with other juveniles and nonbreeders. They remain in these flocks until they form breeding pairs between two and seven years old. As a conspicuous ground-dwelling species, sandhill cranes are at risk from predators, which are probably the main nonhuman source of mortality. Mammals such as
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
es,
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s,
coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
s,
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
,
cougar The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
s,
bobcat The bobcat (''Lynx rufus''), also known as the red lynx, is a medium-sized cat native to North America. It ranges from southern Canada through most of the contiguous United States to Oaxaca in Mexico. It is listed as Least Concern on the IUC ...
s, and
lynx A lynx is a type of wild cat. Lynx may also refer to: Astronomy * Lynx (constellation) * Lynx (Chinese astronomy) * Lynx X-ray Observatory, a NASA-funded mission concept for a next-generation X-ray space observatory Places Canada * Lynx, Ontar ...
hunt them given any opportunity, the first three mainly hunting large numbers of young cranes, the latter four types more rarely taking full-grown cranes in ambush excepting the bobcat.
Corvid Corvidae is a Cosmopolitan distribution, cosmopolitan family (biology), family of oscine passerine birds that contains the crows, ravens, rook (bird), rooks, jackdaws, jays, magpies, treepies, choughs, and nutcracker (bird), nutcrackers. In coll ...
s, such as
raven A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
s and
crow A crow is a bird of the genus ''Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifical ...
s,
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...
s, and smaller raptors such as
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s (largely
northern harrier The northern harrier (''Circus hudsonius''), or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost USA. The northern harrier migrates to more southerly areas ...
s or
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members with ...
s) feed on young cranes and eggs. Cranes of all ages are hunted by both North American species of
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s. Mainly chicks and possibly a few adults may be preyed on by
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air"), or the hoot owl, is a large owl native to the Americas. It is an extrem ...
s, and even a much smaller
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
has successfully killed a adult sandhill crane in a stoop.Walkinshaw, L. H. 1949. ''The Sandhill Cranes''. Cranbrook Inst. Sci. Bull. 29. Bloomfield Hills, MI.Walkinshaw, L. H. 1973. ''Cranes of the world''. Winchester Press, New York. In
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
and
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
, the most serious predators of flighted juveniles and adults has been cited as
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
s and bobcats, the most serious predators of chicks are reportedly
coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
s,
raven A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
s,
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s,
American mink The American mink (''Neogale vison'') is a semiaquatic species of mustelid native to North America, though human intervention has expanded its range to many parts of Europe, Asia and South America. Because of range expansion, the American mink i ...
, and
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air"), or the hoot owl, is a large owl native to the Americas. It is an extrem ...
s, roughly in descending order. In
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, the
American crocodile The American crocodile (''Crocodylus acutus'') is a species of crocodilian found in the Neotropics. It is the most widespread of the four extant species of crocodiles from the Americas, with populations present from South Florida and the coasts ...
and
alligator An alligator is a large reptile in the Crocodilia order in the genus ''Alligator'' of the family Alligatoridae. The two extant species are the American alligator (''A. mississippiensis'') and the Chinese alligator (''A. sinensis''). Additiona ...
can take surprisingly large numbers of sandhill cranes, especially recent fledglings. Sandhill cranes defend themselves and their young from aerial predators by jumping and kicking. Actively brooding adults are more likely to react aggressively to potential predators to defend their chicks than wintering birds, which most often normally try to evade attacks on foot or in flight. For land predators, they move forward, often hissing, with their wings open and bills pointed. If the predator persists, the crane stabs with its bill (which is powerful enough to pierce the skull of a small carnivore) and kicks. File:Grus canadensis -Bosque del Apache National Wildlife Refuge, New Mexico, USA-8.ogv, A huge flock at Bosque del Apache National Wildlife Refuge, New Mexico File:Baby Mississippi Sandhill Crane.JPG, A baby Mississippi sandhill crane is weighed at
White Oak Conservation White Oak Conservation, which is part of Walter Conservation, is a conservation center in northeastern Florida. It has long been dedicated to the conservation and care of endangered and threatened species, including rhinoceros, okapi, Bongo (a ...
. File:Sc2sf.jpg, A human feeding a sandhill crane, a docile and gentle bird


Status and conservation

In the 1930s, sandhill cranes were generally
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
east of the
Mississippi River The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it f ...
, but their populations have recovered, with an estimated 98,000 in the region in 2018, a substantial increase over the previous year. Although sandhill cranes are not considered threatened as a species, the three southernmost subspecies are quite rare. Resident populations, not migratory birds, cannot choose secure breeding habitat. Many
subpopulation In statistics, a population is a set of similar items or events which is of interest for some question or experiment. A statistical population can be a group of existing objects (e.g. the set of all stars within the Milky Way galaxy) or a hypothe ...
s were destroyed by hunting or habitat change. The greater sandhill crane proper initially suffered most; by 1940, probably fewer than 1,000 birds remained. Populations have since increased greatly again. At nearly 100,000, they are still fewer than the lesser sandhill crane, which, at about 400,000 individuals continent-wide, is the most plentiful extant crane. Some migratory populations of sandhill cranes face population threats due to interspecies competition with snow geese. Since the 1990s, snow geese have eaten waste corn on which the cranes also rely prior to migration. Sandhill crane populations are also threatened by hunting. Hunting cranes is legal throughout the states of the Central Flyway, from the Dakotas and Wyoming south to Oklahoma and Texas. Nebraska is the sole state along the Central Flyway where hunting cranes is illegal. Despite losses from hunting, interspecies competition and other pressures such as habitat loss, the species has expanded its range. Since the early 2000s, the sandhill crane has expanded both its winter (nonbreeding) and breeding ranges northward, including into upstate New York. In the 21st century, parts of the
Midwestern United States The Midwestern United States, also referred to as the Midwest or the American Midwest, is one of four census regions of the United States Census Bureau (also known as "Region 2"). It occupies the northern central part of the United States. I ...
have seen an extensive rebound of the species. The transplantation of wild birds and introduction of captive-reared birds into suitable low-population areas have been called viable management techniques. The Mississippi sandhill crane has lost the most range; it used to live along most of the northern
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...
coast, and its range was once nearly
parapatric In parapatric speciation, two subpopulations of a species evolve reproductive isolation from one another while continuing to exchange genes. This mode of speciation has three distinguishing characteristics: 1) mating occurs non-randomly, 2) gene ...
with that of its eastern neighbor. As of 2013, about 25 breeding pairs exist in an intensively managed population. The
Mississippi Sandhill Crane National Wildlife Refuge The Mississippi Sandhill Crane National Wildlife Refuge was established in 1975 to safeguard the endangered Mississippi sandhill crane and its unique disappearing wet pine savanna habitat. The refuge consists of more than in four units and is no ...
—established in 1975 when fewer than 35 of the birds existed—has the biggest release program for cranes on Earth, and 90% of the cranes there were raised in captivity. The second viable egg from a two-egg nest was occasionally removed from the nests, starting in 1965, to become part of a captive flock. This breeding flock is divided between the Audubon Institute's Species Survival Center and
White Oak Conservation White Oak Conservation, which is part of Walter Conservation, is a conservation center in northeastern Florida. It has long been dedicated to the conservation and care of endangered and threatened species, including rhinoceros, okapi, Bongo (a ...
in
Yulee, Florida Yulee is a Census county division (CCD) in Nassau County, Florida, United States. The population is currently 28,798 as of April 2018. Yulee is part of the Jacksonville metropolitan area, which was home to 1,504,980 people in 2017. Today, t ...
. These cranes have produced offspring for annual releases into the refuge. A Mississippi sandhill crane was the first bird to hatch from an egg fertilized by sperm that was thawed from a
cryogenic In physics, cryogenics is the production and behaviour of materials at very low temperatures. The 13th IIR International Congress of Refrigeration (held in Washington DC in 1971) endorsed a universal definition of “cryogenics” and “cr ...
state. This occurred at the
Audubon Institute The Audubon Nature Institute is a family of museums and parks dedicated to nature and based in New Orleans, Louisiana. It consists of the Audubon Zoo, Audubon Aquarium of the Americas, Audubon Louisiana Nature Center, Audubon Park, Woldenberg R ...
, as part of this subspecies' endangered species recovery plan. About 300 Cuban sandhill cranes exist; this is the least known of the populations. In January 2019, 25 to 30 thousand cranes (both greater and lesser subspecies) were found wintering at the Whitewater Draw State Wildlife Area near McNeal in southeast Arizona. Sandhill cranes have been tried as foster parents for
whooping crane The whooping crane (''Grus americana'') is the tallest North American bird, named for its whooping sound. It is an endangered crane species. Along with the sandhill crane (''Antigone canadensis''), it is one of only two crane species native to N ...
s in reintroduction schemes. This failed when the whooping cranes imprinted on their foster parents, later did not recognize other whooping cranes as their
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
s, and unsuccessfully tried to pair with sandhill cranes, instead. File:Sandhill Crane's.jpg, Florida sandhill crane,
Ocala National Forest The Ocala National Forest ls the second largest nationally protected forest in the U.S. State of Florida. It covers of northern Florida. It is located three miles (5 km) east of Ocala and southeast of Gainesville. The Ocala National For ...
File:Sandhill Crane JCB.jpg, Sandhill crane at
Jonathan Dickinson State Park Jonathan Dickinson State Park is a Florida State Park and historic site located in Martin County, Florida, between Hobe Sound and Tequesta. The park includes the Elsa Kimbell Environmental Education and Research Center and a variety of natural ...
, Florida File:Grus canadensis -British Columbia, Canada -upper body-8.jpg, In
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
, Canada


Vagrancy

Sandhill cranes occasionally reach Europe as
vagrants Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
. The first British record was on
Fair Isle Fair Isle (; sco, Fair Isle; non, Friðarey; gd, Fara) is an island in Shetland, in northern Scotland. It lies about halfway between mainland Shetland and Orkney. It is known for its bird observatory and a traditional style of knitting. Th ...
in April 1981, and the second was in
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
in 1991. Small groups have also been seen in parts of eastern China and
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
. In 2022, reports emerged of regular sightings of sandhill cranes in New Brunswick, on the Atlantic coast of Canada.


See also

* Cranes in Britain *
Grulla National Wildlife Refuge Grulla National Wildlife Refuge is located primarily in eastern New Mexico in Roosevelt County, southwest of the intersection of State Highway 88 and the Texas - New Mexico border about 25 miles southeast of Portales, New Mexico and southeast of ...
*
Muleshoe National Wildlife Refuge Muleshoe National Wildlife Refuge is a wildlife refuge located about south of Muleshoe, Texas, on Texas State Highway 214. It is the oldest national wildlife refuge in Texas, having been established as the Muleshoe Migratory Waterfowl Refuge ...


References


Further reading

* Meine, Curt D. & Archibald, George W. (eds.) (1996)
"Sandhill Crane (''Grus canadensis'')"
In: The cranes: Status survey and conservation action plan. IUCN, Gland, Switzerland, and Cambridge, U.K.


External links


International Crane Foundation's Sandhill Crane page

Audio file of cranes "unison calling"
at savingcranes.org
Sandhill Crane Bird Sound
at Florida Museum of Natural History

– Cornell Lab of Ornithology

– USGS Patuxent Bird Identification InfoCenter


National Audubon Society, Rowe Sanctuary, Nebraska


– Viewing Sites Info & Maps, Photos, Video


Mississippi Sandhill Crane National Wildlife Refuge
*

(for
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
,
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
) with Range Map at bird-stamps.com
Sandhill Crane Migrations Revealed by Satellites
Documentary produced by
Oregon Field Guide ''Oregon Field Guide'' is a weekly television program produced by Oregon Public Broadcasting focusing on recreation, the outdoors, and environmental issues in the state of Oregon. The show has become part of the Oregon zeitgeist. Steve Amen is th ...

Sandhill cranes breeding and biology in Alaska
at christyyuncker.com *
''Música de las Grullas: Una historia natural de las grullas de América''
(2014) by
Paul Johnsgard Paul Austin Johnsgard (28 June 1931 – 28 May 2021) was an ornithologist, artist and emeritus professor at the University of Nebraska. His works include nearly fifty books including several monographs, principally about the waterfowl and cranes. ...
, Enrique Weir, & Karine Gil-Weir {{Authority control
sandhill crane The sandhill crane (''Antigone canadensis'') is a species of large crane of North America and extreme northeastern Siberia. The common name of this bird refers to habitat like that at the Platte River, on the edge of Nebraska's Sandhills on t ...
Birds of North America Birds of Canada Birds of Cuba Birds of the Caribbean Native birds of the Western United States Native birds of Alaska Wading birds
sandhill crane The sandhill crane (''Antigone canadensis'') is a species of large crane of North America and extreme northeastern Siberia. The common name of this bird refers to habitat like that at the Platte River, on the edge of Nebraska's Sandhills on t ...
Articles containing video clips Taxa named by Carl Linnaeus Taxobox binomials not recognized by IUCN