The greater mouse-eared bat (''Myotis myotis'') is a

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

an species of

bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most bi ...

in the family

Vespertilionidae Vespertilionidae is a family of microbats, of the order Chiroptera, flying, insect-eating mammals variously described as the common, vesper, or simple nosed bats. The vespertilionid family is the most diverse and widely distributed of bat familie ...

Description

''Myotis myotis'' is a large bat with a long, broad muzzle and big, long ears. The body's dorsal side is brown to reddish-brown, while the ventral side is dirty white or beige. The tragus forms half of the ear, with a small black tip in most individuals. Wing membranes are brownish in colour. The Greater mouse-eared bat is relatively large for a member of the genus ''Myotis'', weighing up to and measuring 8 to 9 cm from head to tail (a little larger than a

house mouse The house mouse (''Mus musculus'') is a small mammal of the order Rodentia, characteristically having a pointed snout, large rounded ears, and a long and almost hairless tail. It is one of the most abundant species of the genus '' Mus''. Althoug ...

, ''Mus musculus''), making it one of the largest European bats. It has a 40 cm wingspan, with a forearm length of 6 cm, and a 4 to 5 cm long tail. The average lifespan of a greater mouse-eared bat is 3–4 years, although particular individuals have lived up to 14 years.

Distribution

The greater mouse-eared bat can be found throughout Europe, with populations in most European countries except

Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...

Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...

Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...

Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...

and the

Scandinavian Peninsula The Scandinavian Peninsula ( sv, Skandinaviska halvön; no, Den skandinaviske halvøy (Bokmål) or nn, Den skandinaviske halvøya; fi, Skandinavian niemimaa) is a peninsula located in Northern Europe, which roughly comprises the mainlands ...

. It is also found on many Mediterranean islands, such as

Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...

Malta Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...

, and the

Gymnesian Islands The Gymnesians ( ca, Illes Gimnèsies , es, Gimnesias ), or Gymnesic Islands ( ca, Illes Gimnèsiques), is a collective name given to the two largest (and easternmost) Balearic Islands, Mallorca and Menorca, distinguishing them from the Pityu ...

. In the Middle East, ''Myotis myotis'' has been found in

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...

Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...

Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...

and

Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus li ...

. It possibly lives in the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

. In the

Balearics The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...

, ''Myotis myotis'' can be found on the islands of

Mallorca Mallorca, or Majorca, is the largest island in the Balearic Islands, which are part of Spain and located in the Mediterranean. The capital of the island, Palma, is also the capital of the autonomous community of the Balearic Islands. The Bal ...

and

Menorca Menorca or Minorca (from la, Insula Minor, , smaller island, later ''Minorica'') is one of the Balearic Islands located in the Mediterranean Sea belonging to Spain. Its name derives from its size, contrasting it with nearby Majorca. Its capi ...

. It is the most abundant species of bat on Mallorca, with several caves containing large colonies of 200, 400 and 500 individuals. It is also very common in the

Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...

and

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

. During the 20th century, this species was very rare in Great Britain, occurring only in southern

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

. However, the bats at the only known

hibernation Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...

roost declined until only a few males were left, and when these disappeared the species was believed locally extinct. However, occasional individuals have been discovered in recent years, suggesting either that a colony survives or that further animals have colonized Great Britain from mainland Europe. In 2012, a LIFE-Nature project was initiated, aiming to protect the several thousands of greater mouse-eared bat in the Gola della Rossa and Frasassi Nature Park in

the Marches Marche ( , ) is one of the twenty regions of Italy. In English, the region is sometimes referred to as The Marches ( ). The region is located in the central area of the country, bordered by Emilia-Romagna and the republic of San Marino to the ...

Behaviour

Like its relatives, the greater mouse-eared bat is an

insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...

, feeding on various

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

s; however, unlike many bats, it does not capture its prey by using echolocation in flight. Instead, it gleans it from the ground, locating prey passively by listening for the noises produced by insects such as carabid beetles, other

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

s and large

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

s, as well as

centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...

s and

spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...

s. As a result, it uses echolocation only for spatial orientation, even if it emits ultrasound calls when approaching prey. ''Myotis myotis'' forages in open deciduous woodlands, forest edges and pastures. It roosts underground throughout the year, although roosts in Northern Europe are also located in attics and lofts of buildings. More rarely, small tree-based colonies also occur. Colonies on Mediterranean islands such as Mallorca usually have up to 500 individuals, though colonies of over 4.500 bats have been found in continental Europe. It has been known to roost with other bats, such as ''Myotis capaccinii'' and ''Miniopterus schreibersi''. In mainland Europe, ''Myotis myotis'' performs annual dispersions of up to 200 km in spring, although most are only of 10 km or so. As with most

vespertilionid Vespertilionidae is a family of microbats, of the order Chiroptera, flying, insect-eating mammals variously described as the common, vesper, or simple nosed bats. The vespertilionid family is the most diverse and widely distributed of bat familie ...

bats, the greater mouse-eared bat exhibits slight

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...

, with females being slightly larger than males. Mating takes place in autumn, with females giving birth from May to June. Each female has 1–2 pups, which are carried around for about 45 days. During this period, females form nursery colonies from which males are excluded. The frequencies used by this bat species for echolocation lie between 22 and 86 kHz, have most energy at 37 kHz and have an average duration of 6.0 ms.

References

{{Taxonbar, from=Q723413 Mouse-eared bats Bats of Asia Mammals of Turkey Fauna of the Middle East Mammals of the Middle East Bats of Europe Mammals described in 1797