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The grass carp (''Ctenopharyngodon idella'') is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of large
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
freshwater fish in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest v ...
, native to the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
Far East The ''Far East'' was a European term to refer to the geographical regions that includes East and Southeast Asia as well as the Russian Far East to a lesser extent. South Asia is sometimes also included for economic and cultural reasons. The te ...
, with a native range stretching from northern Vietnam to the
Amur River The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's tenth longest river, forming the border between the Russian Far East and Northeastern China ( Inner Manchuria). The Amur proper is long ...
on the Sino-Russian border.Mandrak and Cudmore. 2004
''Biological Synopsis of Grass Carp (Ctenopharyngodon idella)''
.
This Asian carp is the only species of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Ctenopharyngodon''. Grass carp are resident fish of large turbid
river A river is a natural flowing watercourse, usually freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the ...
s and associated floodplain lakes/
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
s with a wide range of temperature tolerance, and spawn at temperatures of . It is cultivated as a food fish in China for centuries, but was introduced in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
and the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
for aquatic weed control, becoming the fish species with the largest reported farmed production globally, over five million tonnes per year.World aquaculture production of fish, crustaceans, mollusks, etc., by principal species in 2013
FAO Yearbook of Fisheries Statistics 2014


Appearance and anatomy

Grass carp have elongated, chubby, torpedo-shaped body forms. The terminal mouth is slightly oblique with non-fleshy, firm lips, and no barbels.''Grass carp''
in ''fishbase.org''.
The complete lateral line contains 40 to 42 scales. Broad, ridged pharyngeal teeth are arranged in a "2, 4-4, 2" formula. The dorsal fin has eight to 10 soft rays, and the anal fin is set closer to the tail than most cyprinids. Body color is dark olive, shading to brownish-yellow on the sides, with a white belly and large, slightly outlined scales. Grass carp grow very rapidly. Young fish stocked in the spring at will reach over by fall. The typical length is about . The maximum length is and they grow to .


Ecology

Grass carp inhabit lakes,
pond A pond is an area filled with water, either natural or Artificiality, artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% Aquatic plant, emergent vegetation helps in disting ...
s, pools and
backwater Backwater or Backwaters may refer to: Music * ''Backwaters'' (album), a 1982 album by American guitarist Tony Rice * Backwater (band), a jazz fusion band from Mobile, Alabama, or this band's 1976 debut album * "Backwater", a song by Brian Eno fro ...
s of large rivers, preferring large, slow-flowing or standing water bodies with abundant vegetation. In the wild, grass carp spawn in fast-moving rivers, and their eggs, which are slightly heavier than water, develop while drifting downstream, kept in suspension by turbulence. Grass carp require long rivers for the survival of the eggs and very young fish, and the eggs are thought to die if they sink to the bottom. Adults grass carp feed primarily on
aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments ( saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
s, both higher aquatic plants and submerged terrestrial vegetation, but may also eat
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commu ...
,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
s and other
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s. They eat up to three times their own body weight daily, and thrive in small lakes and backwaters that provide an abundant supply of vegetation. According to one study, grass carp live 5-9 years, with the oldest surviving 11 years. In
Silver Lake Silver is a chemical element with the symbol Ag (from the Latin ', derived from the Proto-Indo-European ''h₂erǵ'': "shiny" or "white") and atomic number 47. A soft, white, lustrous transition metal, it exhibits the highest electrical ...
, Washington, a thriving population of grass carp is passing the 15-year mark.


Introduced species

Grass carp have been introduced to many countries around the world. In the Northern Hemisphere, countries and territories of introduction include Japan, the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
,
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
,
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
,
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkm ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
, the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
,
Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
, Sweden,
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establishe ...
, the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...
,
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
, the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
, Switzerland,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
, the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. Th ...
,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the ...
,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central, Eastern, and Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, Serbia to the southwest, Moldova to the east, a ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
,
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, and ...
,
Serbia Serbia (, ; Serbian: , , ), officially the Republic of Serbia (Serbian: , , ), is a landlocked country in Southeastern and Central Europe, situated at the crossroads of the Pannonian Basin and the Balkans. It shares land borders with Hung ...
,
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = ...
,
Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and ...
and
Macedonia Macedonia most commonly refers to: * North Macedonia, a country in southeastern Europe, known until 2019 as the Republic of Macedonia * Macedonia (ancient kingdom), a kingdom in Greek antiquity * Macedonia (Greece), a traditional geographic reg ...
. In the Southern Hemisphere, they have been introduced to
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
,
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
, Australia,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 List of islands of New Zealand, smaller islands. It is the ...
, Fiji and
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
. Grass carp are known to have spawned and established self-reproducing populations in only six of the many larger Northern Hemisphere rivers into which they have been stocked. Their failure to establish populations in other rivers suggests they have quite specific reproductive requirements. In the United States, the species was first imported in 1963 from
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the northe ...
and Malaysia to aquaculture facilities in
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
and
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the ...
. The first release is believed to have been an accidental escape in 1966 from the U.S. Fish and Wildlife Service's Fish Farming Experimental Station in Stuttgart, Arkansas, followed by planned introductions beginning in 1969. Subsequently, authorized, illegal and accidental introductions have been widespread; by the 1970s, the species had been introduced to 40 states, and it has since been reported in 45 of the country's 50 states. In 2013, it was determined to be reproducing in the Great Lakes Basin. It is still stocked in many states as an effective biocontrol for undesirable aquatic vegetation, many species of which are themselves introduced.


Use as weed control

Grass carp were introduced into New Zealand in 1966 to control the growth of aquatic plants. Unlike the other introduced fish brought to New Zealand, the potential value and impact of grass carp was investigated in secure facilities prior to their use in field trials. They are now approved by the New Zealand government for aquatic weed control, although each instance requires specific authorization."Grass carp for weed control"
DOC
In the Netherlands, the species was also introduced in 1973 to control over-abundant aquatic weeds. The release was controlled and regulated by the Dutch Ministry of Agriculture, Nature, and Food Quality. In both of these countries, control is made easier because grass carp are very unlikely to naturally reproduce because of their very specific breeding requirements, but elsewhere, control is obtained by the use of sterile, triploid fish.


Fishing for grass carp

Grass carp grow large and are strong fighters when hooked on a line, but because of their vegetarian habits and their wariness, they can be difficult to catch via
angling Angling is a fishing technique that uses a fish hook or "angle" (from Old English ''angol'') attached to a fishing line to tether individual fish in the mouth. The fishing line is usually manipulated via a fishing rod, although rodless techni ...
. The IGFA World record for a grass carp caught on line and hook is , caught in
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Mac ...
in 2009."Grass carp"
IGFA Online. Retrieved 22 June 2016.
The fish are also popular sport fish in areas where bowfishing is legal. Where grass carp populations are maintained through
stocking Stockings (also known as hose, especially in a historical context) are close-fitting, variously elastic garments covering the leg from the foot up to the knee or possibly part or all of the thigh. Stockings vary in color, design, and transpare ...
as a biocontrol for noxious weeds, fishermen are typically asked to return any caught to the water alive and unharmed.


References


External links


Species Profile - Grass Carp (''Ctenopharyngodon idella'')
National Invasive Species Information Center,
United States National Agricultural Library The United States National Agricultural Library (NAL) is one of the world's largest agricultural research libraries, and serves as a national library of the United States and as the library of the United States Department of Agriculture. Located ...
. Lists general information and resources for Grass Carp. * * {{taxonbar, from=Q76098 grass carp Commercial fish Freshwater fish of China Fish of East Asia Herbivorous vertebrates Taxa named by Achille Valenciennes grass carp Squaliobarbinae Chinese seafood dishes