Graphium Antiphates
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''Graphium antiphates'', the five-bar swordtail, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of papilionid
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
found in
south South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. The species was first described by
Pieter Cramer Pieter Cramer (21 May 1721 (baptized) – 28 September 1776), was a wealthy Dutch merchant in linen and Spanish wool, remembered as an entomologist. Cramer was the director of the Zealand Society, a scientific society located in Flushing, and a mem ...
in 1775.


Description

The ground colour of the upperside of both males and females is white. The forewing has the cell crossed by five short black bands, of which the basal extends to the dorsum, the sub-basal into interspace 1, the medial and pre-apical up to the median vein, and the apical or fifth along the discocellulars; this last band extends broadly on both sides of the veinlets and terminates at the lower apex of the cell; beyond these are broad postdiscal and terminal black transverse bands from costa to tornal angle; the two bands coalesce below vein 4 and terminate in a point at the tornus; the white portions of the cell anteriorly overlaid with pale green; short broken glossy green bands between the black cellular apical band and the discal band and anteriorly between the latter and the terminal band. The upperside of the hindwing has the basal three-fourths uniformly white, with black markings on the underside that show through; the terminal fourth dark grey traversed by a curved irregular subterminal series of black crescent shaped marks that ends in a black tornal spot and a terminal black band that follows the indentations of the wing; the emargination (notches in a margin) below the black tornal spot are edged with ochraceous; the tail blackish grey, edged and tipped with white. The underside of the forewing is similar to the upperside in markings but the green shading over the white portions in the basal half of the cell more decided; the discal and terminal transverse black bands are separate, and are not joined posteriorly, the former edged posteriorly on both sides by dark grey due to the black on the upperside that shows through by transparency. The underside of the hindwing is half green on the basal part while the outer half white; a large black tornal spot; a black line along the dorsum that curves above the tornal spot outwards to vein 2; a straight subbasal black band from costa across cell that terminates on vein 2, where it joins the dorsal black line; a broader black band from costa across apex of cell extended into base of interspace 3; an irregular discal series of black markings curved inwards posteriorly towards the tornal spot; a subterminal series of very small slender black lunules in pairs, the ground colour on the inner side of these darkened to rich ochreous yellow; lastly, a series of short terminal black bars in the interspaces so arranged as to follow indentations of the termen; tail dusky black edged with white. Antenna black; head and thorax anteriorly with a broad black medial band, rest of thorax bluish; abdomen white, marked beneath on each side by a black stripe.
Race Race, RACE or "The Race" may refer to: * Race (biology), an informal taxonomic classification within a species, generally within a sub-species * Race (human categorization), classification of humans into groups based on physical traits, and/or s ...
''alcibiades'', Fabr. is the most widely spread race of ''antiphates'', from which it differs as follows: Upperside of males and females, all the black markings shorter and narrower. Forewing: the discal and terminal bands separate, the former rarely extended below vein 3, the latter in no specimen reaches the dorsal margin. Hindwing: the broad grey area on the terminal margin reduced to a small patch of grey at the apices of interspaces 2 to 4; the subterminal black markings rarely present anteriorly, generally confined to the limits of the grey patch. Underside: the extent of the black markings similarly reduced, otherwise as in the typical form. The width and length of the transverse black markings on the upperside of the forewing, also the extent of the grey terminal area and the presence or absence of the black subterminal markings on the upperside of the hindwing, are all very variable.
Form Form is the shape, visual appearance, or configuration of an object. In a wider sense, the form is the way something happens. Form also refers to: *Form (document), a document (printed or electronic) with spaces in which to write or enter data ...
''nebulosus'', Butler, is a
melanistic The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair. Pseudomelanism, also called abundism, is another variant of pi ...
variety recorded from Sikkim. Var. ''continentalis'', Eimer, has the caudal area of the hindwing on the upperside suffused with black. Var. ''itamputi'', Butler, has the postdiscal and terminal black bands on the upperside of the forewing united posteriorly, but neither band extends up to the tornus. Var. ''ceylanicus'', Eimer, has the basal two bands on the upperside of the forewing extended beyond the median vein, the preapical cellular band not triangular and extended to the median vein.


Habits

It is known to
mud-puddle Mud-puddling, or simply puddling, is a behaviour most conspicuous in butterflies, but occurs in other animals as well, mainly insects; they seek out nutrients in certain moist substances such as rotting plant matter, mud and carrion and they suck ...
.


Life history

The caterpillar is white in the early stage and turns yellow in the last
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or ass ...
. In its late stage, it is like the caterpillar of ''
Pathysa nomius ''Graphium nomius'', the spot swordtail, is a butterfly found in South and Southeast Asia that belongs to the swallowtail family. The species was first described by Eugenius Johann Christoph Esper in 1793. One of the grandest sights is a host o ...
''. (Davidson & Aitken quoted in Bingham) The green pupa, is as in all swallowtails, is held by a silk girdle. Said to be found mainly on the plant '' Unona lawii'' and never under stones. (Davidson and Aitken)


Gallery

File:FivebarSwordtail-Aralam-DSC 3051A.jpg, Five-bar swordtails in
Aralam Wildlife Sanctuary Aralam Wildlife Sanctuary is the northernmost wildlife sanctuary of Kerala, southwest India. It is in area and located on the western slope of the Western Ghats. It was established in 1984. The headquarters of the sanctuary is near Iritty ...
, India File:Graphium antiphates mudpuddling someshwara.JPG, Five-bar swordtails
mud-puddling Mud-puddling, or simply puddling, is a behaviour most conspicuous in butterflies, but occurs in other animals as well, mainly insects; they seek out nutrients in certain moist substances such as rotting plant matter, mud and carrion and they suck ...
in
Someshwara Wildlife Sanctuary Someshwara Wildlife Sanctuary is a protected wildlife sanctuary in the Western Ghats of Karnataka state in India. It is named after the presiding deity "Lord Someshwara" of the famed Someshwara temple located within the sanctuary. The sanctuary ...
, India File:PathysaAlcibiadesLP_474.jpg, Larva and pupa


References

{{Taxonbar, from=Q775758 Pathysa
antiphates In Greek mythology, Antiphates (; Ancient Greek: Ἀντιφάτης) is the name of five characters. * Antiphatês, son of Melampus and Iphianeira, the daughter of Megapenthes. He married Zeuxippe, the daughter of Hippocoon. Their children were O ...
Butterflies of Asia Fauna of Pakistan Butterflies of Singapore Butterflies of Indochina Taxa named by Pieter Cramer Butterflies described in 1775