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A grape is a fruit, botanically a berry, of the deciduous woody
vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themselv ...
s of the flowering plant genus ''
Vitis ''Vitis'' (grapevine) is a genus of 79 accepted species of vining plants in the flowering plant family Vitaceae. The genus is made up of species predominantly from the Northern Hemisphere. It is economically important as the source of grapes, ...
''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years ago, and the fruit has been used as human food over history. Eaten fresh or in dried form (as
raisin A raisin is a dried grape. Raisins are produced in many regions of the world and may be eaten raw or used in cooking, baking, and brewing. In the United Kingdom, Ireland, New Zealand, and Australia, the word ''raisin'' is reserved for the ...
s, currants and sultanas), grapes also hold cultural significance in many parts of the world, particularly for their role in
winemaking Winemaking or vinification is the production of wine, starting with the selection of the fruit, its fermentation into alcohol, and the bottling of the finished liquid. The history of wine-making stretches over millennia. The science of wine and ...
. Other grape-derived products include various types of jam, juice, vinegar and oil.


History

The
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
is generally described as the homeland of grape and the cultivation of this plant began there 6,000–8,000 years ago.
Yeast Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom. The first yeast originated hundreds of millions of years ago, and at least 1,500 species are currently recognized. They are estimated to constitut ...
, one of the earliest domesticated
microorganism A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
s, occurs naturally on the skins of grapes, leading to the discovery of alcoholic drinks such as wine. The earliest archeological evidence for a dominant position of wine-making in human culture dates from 8,000 years ago in
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
. The oldest known winery was found in
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ''Ox ...
, dating to around 4000 BC. By the 9th century AD, the city of
Shiraz Shiraz (; fa, شیراز, Širâz ) is the List of largest cities of Iran, fifth-most-populous city of Iran and the capital of Fars province, Fars Province, which has been historically known as Pars (Sasanian province), Pars () and Persis. As o ...
was known to produce some of the finest wines in the Middle East. Thus it has been proposed that Syrah red wine is named after
Shiraz Shiraz (; fa, شیراز, Širâz ) is the List of largest cities of Iran, fifth-most-populous city of Iran and the capital of Fars province, Fars Province, which has been historically known as Pars (Sasanian province), Pars () and Persis. As o ...
, a city in Persia where the grape was used to make
Shirazi wine Shiraz wine refers to two different wines. Historically, the name refers to the wine produced around the city of Shiraz in present-day Iran.Entry on ''"Persia"'' in J. Robinson (ed), ''"The Oxford Companion to Wine"'', Third Edition, p. 512-513, ...
.Hugh Johnson, ''"The Story of Wine"'', New Illustrated Edition, p. 58 & p. 131, Mitchell Beazley 2004, Ancient Egyptian
hieroglyphics Egyptian hieroglyphs (, ) were the formal writing system used in Ancient Egypt, used for writing the Egyptian language. Hieroglyphs combined logographic, syllabic and alphabetic elements, with some 1,000 distinct characters.There were about 1,00 ...
record the cultivation of purple grapes, and history attests to the ancient
Greeks The Greeks or Hellenes (; el, Έλληνες, ''Éllines'' ) are an ethnic group and nation indigenous to the Eastern Mediterranean and the Black Sea regions, namely Greece, Cyprus, Albania, Italy, Turkey, Egypt, and, to a lesser extent, oth ...
, Cypriots,
Phoenicians Phoenicia () was an ancient Semitic-speaking peoples, ancient thalassocracy, thalassocratic civilization originating in the Levant region of the eastern Mediterranean, primarily located in modern Lebanon. The territory of the Phoenician city-st ...
, and
Romans Roman or Romans most often refers to: *Rome, the capital city of Italy * Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a lette ...
growing purple grapes both for eating and wine production. The growing of grapes would later spread to other regions in Europe, as well as North Africa, and eventually in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. In 2005 a team of archaeologists concluded that some
Chalcolithic The Copper Age, also called the Chalcolithic (; from grc-gre, χαλκός ''khalkós'', "copper" and  ''líthos'', "stone") or (A)eneolithic (from Latin '' aeneus'' "of copper"), is an archaeological period characterized by regular ...
wine jars, which were discovered in Cyprus in the 1930s, were the oldest of their kind in the world, dating back to 3,500 BC. Moreover, Commandaria, a sweet dessert wine from Cyprus, is the oldest manufactured wine in the world, its origins traced as far back as 2000 BC. In North America, native grapes belonging to various species of the genus ''Vitis'' proliferate in the wild across the continent, and were a part of the diet of many Native Americans, but were considered by early European colonists to be unsuitable for wine. In the 19th century,
Ephraim Bull Ephraim Wales Bull (March 4, 1806 – September 26, 1895) was an American farmer, best known for the creation of the Concord grape. Biography Ephraim Wales Bull was born on March 4, 1806, in Boston, Massachusetts.
of
Concord, Massachusetts Concord () is a town in Middlesex County, Massachusetts, in the United States. At the 2020 census, the town population was 18,491. The United States Census Bureau considers Concord part of Greater Boston. The town center is near where the confl ...
, cultivated seeds from wild ''
Vitis labrusca ''Vitis labrusca'', the fox grape, is a species of grapevines belonging to the ''Vitis'' genus in the flowering plant family Vitaceae. The vines are native to eastern North America and are the source of many grape cultivars, including Catawba, C ...
'' vines to create the
Concord grape The Concord grape is a cultivar derived from the grape species '' Vitis labrusca'' (also known as fox grape) that are used as table grapes, wine grapes and juice grapes. They are often used to make grape jelly, grape juice, grape pies, grape-fl ...
which would become an important agricultural crop in the United States.Jancis Robinson, ''Vines, Grapes & Wines'' (Mitchell Beazley, 1986, ), pp 8, 18, 228.


Description

Grapes are a type of
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
that grow in clusters of 15 to 300, and can be crimson, black, dark blue, yellow, green, orange, and pink. "White" grapes are actually green in color, and are evolutionarily derived from the purple grape.
Mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA replication, DNA or viral repl ...
s in two
regulatory genes A regulator gene, regulator, or regulatory gene is a gene involved in controlling the expression of one or more other genes. Regulatory sequences, which encode regulatory genes, are often at the five prime end (5') to the start site of transcript ...
of white grapes turn off production of anthocyanins, which are responsible for the color of purple grapes. Anthocyanins and other
pigment A pigment is a colored material that is completely or nearly insoluble in water. In contrast, dyes are typically soluble, at least at some stage in their use. Generally dyes are often organic compounds whereas pigments are often inorganic compo ...
chemicals of the larger family of
polyphenols Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some of ...
in purple grapes are responsible for the varying shades of purple in red wines. Grapes are typically an ellipsoid shape resembling a
prolate spheroid A spheroid, also known as an ellipsoid of revolution or rotational ellipsoid, is a quadric surface obtained by rotating an ellipse about one of its principal axes; in other words, an ellipsoid with two equal semi-diameters. A spheroid has ci ...
.


Nutrition

Raw grapes are 81% water, 18%
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s, 1%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
, and have negligible
fat In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food. The term often refers specifically to triglycerides (triple est ...
(table). A reference amount of raw grapes supplies of
food energy Food energy is chemical energy that animals (including humans) derive from their food to sustain their metabolism, including their muscle, muscular activity. Most animals derive most of their energy from aerobic respiration, namely combining the ...
and a moderate amount of
vitamin K Vitamin K refers to structurally similar, fat-soluble vitamers found in foods and marketed as dietary supplements. The human body requires vitamin K for post-synthesis modification of certain proteins that are required for blood coagulation ...
(14% of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of healthy ...
), with no other micronutrients in significant content.


Grapevines

Most domesticated grapes come from cultivars of ''
Vitis vinifera ''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran. There are curre ...
'', a grapevine native to the Mediterranean and Central Asia. Minor amounts of fruit and wine come from American and Asian species such as: * ''
Vitis amurensis ''Vitis amurensis'', the Amur grape, is a species of grape native to the Asian continent. Its name comes from the Amur Valley in Russia and China. It is very resistant to frost, but is not tolerant to drought. Selections vary, but as a speci ...
'', the most important Asian species * ''
Vitis labrusca ''Vitis labrusca'', the fox grape, is a species of grapevines belonging to the ''Vitis'' genus in the flowering plant family Vitaceae. The vines are native to eastern North America and are the source of many grape cultivars, including Catawba, C ...
'', the North American table and grape juice grapevines (including the
Concord Concord may refer to: Meaning "agreement" * Pact or treaty, frequently between nations (indicating a condition of harmony) * Harmony, in music * Agreement (linguistics), a change in the form of a word depending on grammatical features of other ...
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
), sometimes used for wine, are native to the Eastern United States and Canada. * '' Vitis mustangensis'' (the mustang grape), found in Mississippi, Alabama, Louisiana, Texas, and Oklahoma * ''
Vitis riparia ''Vitis riparia'' Michx, with common names riverbank grape or frost grape, is a vine indigenous to North America. As a climbing or trailing vine, it is widely distributed across central and eastern Canada and the central and northeastern parts ...
'', a wild vine of North America, is sometimes used for winemaking and for jam. It is native to the entire Eastern United States and north to
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
. * ''
Vitis rotundifolia ''Vitis rotundifolia'', or muscadine, is a grapevine species native to the southeastern and south-central United States. The growth range extends from Florida to New Jersey coast, and west to eastern Texas and Oklahoma. It has been extensively ...
'' (the muscadine), used for jams and wine, is native to the Southeastern United States from
Delaware Delaware ( ) is a state in the Mid-Atlantic region of the United States, bordering Maryland to its south and west; Pennsylvania to its north; and New Jersey and the Atlantic Ocean to its east. The state takes its name from the adjacent Del ...
to the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...
.


Trade


Distribution and production

According to the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
(FAO), 75,866 square kilometers of the world are dedicated to grapes. Approximately 71% of world grape production is used for wine, 27% as fresh fruit, and 2% as
dried fruit Dried fruit is fruit from which the majority of the original water content has been removed either naturally, through sun drying, or through the use of specialized dryers or dehydrators. Dried fruit has a long tradition of use dating back to th ...
. A portion of grape production goes to producing grape juice to be reconstituted for fruits canned "with no added sugar" and "100% natural". The area dedicated to vineyards is increasing by about 2% per year. There are no reliable statistics that break down grape production by variety. It is believed that the most widely planted variety is Sultana, also known as Thompson Seedless, with at least 3,600 km2 (880,000 acres) dedicated to it. The second most common variety is
Airén Airén is a variety of ''Vitis vinifera'', a white grape commonly used in winemaking. This grape is native to Spain where it represents almost a quarter of all grapes grown. As of 2010, Airén was estimated to be the world's 3rd most grown grape ...
. Other popular varieties include Cabernet Sauvignon,
Sauvignon blanc is a green-skinned grape variety that originates from the Bordeaux region of France. The grape most likely gets its name from the French words ''sauvage'' ("wild") and ''blanc'' ("white") due to its early origins as an indigenous grape in ...
, Cabernet Franc, Merlot, Grenache,
Tempranillo Tempranillo (also known as Ull de Llebre, Cencibel, Tinto Fino and Tinta del Pais in Spain, Aragonez or Tinta Roriz in Portugal, and several other synonyms elsewhere) is a black grape variety widely grown to make full-bodied red wines in its ...
,
Riesling Riesling (, ; ) is a white grape variety that originated in the Rhine region. Riesling is an aromatic grape variety displaying flowery, almost perfumed, aromas as well as high acidity. It is used to make dry, semi-sweet, sweet, and sparkling wh ...
, and
Chardonnay Chardonnay (, , ) is a green-skinned grape variety used in the production of white wine. The variety originated in the Burgundy wine region of eastern French wine, France, but is now grown wherever wine is produced, from English wine, Englan ...
.


Table and wine grapes

Commercially cultivated grapes can usually be classified as either
table Table may refer to: * Table (furniture), a piece of furniture with a flat surface and one or more legs * Table (landform), a flat area of land * Table (information), a data arrangement with rows and columns * Table (database), how the table data ...
or wine grapes, based on their intended method of consumption: eaten raw (table grapes) or used to make
wine Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...
(wine grapes). While almost all of them belong to the same species, ''
Vitis vinifera ''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran. There are curre ...
'', table and wine grapes have significant differences, brought about through
selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant m ...
. Table grape cultivars tend to have large, seedless fruit (see below) with relatively thin skin. Wine grapes are smaller, usually seeded, and have relatively thick skins (a desirable characteristic in winemaking, since much of the aroma in wine comes from the skin). Wine grapes also tend to be very sweet: they are harvested at the time when their juice is approximately 24% sugar by weight. By comparison, commercially produced "100% grape juice", made from table grapes, is usually around 15% sugar by weight.


Seedless grapes

Seedless cultivars now make up the overwhelming majority of table grape plantings. Because grapevines are vegetatively propagated by cuttings, the lack of seeds does not present a problem for reproduction. It is an issue for breeders, who must either use a seeded variety as the female parent or rescue embryos early in development using
tissue culture Tissue culture is the growth of tissues or cells in an artificial medium separate from the parent organism. This technique is also called micropropagation. This is typically facilitated via use of a liquid, semi-solid, or solid growth medium, su ...
techniques. There are several sources of the seedlessness trait, and essentially all commercial cultivators get it from one of three sources:
Thompson Seedless The sultana is a "white" (pale green), oval seedless grape variety also called the sultanina, Thompson Seedless (United States), Lady de Coverly (England), and oval-fruited Kishmish (Iraq, Iran, Israel, Palestine, Pakistan, Afghanistan, India) ...
,
Russian Seedless Russian(s) refers to anything related to Russia, including: *Russians (, ''russkiye''), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *Rossiyane (), Russian language term for all citizens and peo ...
, and
Black Monukka Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have ...
, all being cultivars of ''
Vitis vinifera ''Vitis vinifera'', the common grape vine, is a species of flowering plant, native to the Mediterranean region, Central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran. There are curre ...
''. There are currently more than a dozen varieties of seedless grapes. Several, such as Einset Seedless, Benjamin Gunnels's Prime seedless grapes, Reliance, and Venus, have been specifically cultivated for hardiness and quality in the relatively cold climates of northeastern United States and southern
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
. An offset to the improved eating quality of seedlessness is the loss of potential health benefits provided by the enriched phytochemical content of grape seeds (see Health claims, below).


Raisins, currants and sultanas

In most of Europe and North America, dried grapes are referred to as "raisins" or the local equivalent. In the UK, three different varieties are recognized, forcing the EU to use the term "dried vine fruit" in official documents. A ''
raisin A raisin is a dried grape. Raisins are produced in many regions of the world and may be eaten raw or used in cooking, baking, and brewing. In the United Kingdom, Ireland, New Zealand, and Australia, the word ''raisin'' is reserved for the ...
'' is any dried grape. While ''raisin'' is a French
loanword A loanword (also loan word or loan-word) is a word at least partly assimilated from one language (the donor language) into another language. This is in contrast to cognates, which are words in two or more languages that are similar because th ...
, the word in French refers to the fresh fruit; ''grappe'' (from which the English ''grape'' is derived) refers to the bunch (as in ''une grappe de raisins''). A raisin in French is called ''raisin sec'' ("dry grape"). A '' currant'' is a dried
Zante Zakynthos (also spelled Zakinthos; el, Ζάκυνθος, Zákynthos ; it, Zacinto ) or Zante (, , ; el, Τζάντε, Tzánte ; from the Venetian form) is a Greek island in the Ionian Sea. It is the third largest of the Ionian Islands. Za ...
Black Corinth grape, the name being a corruption of the French ''raisin de Corinthe'' (
Corinth Corinth ( ; el, Κόρινθος, Kórinthos, ) is the successor to an ancient city, and is a former municipality in Corinthia, Peloponnese, which is located in south-central Greece. Since the 2011 local government reform, it has been part o ...
grape). The names of the black and red currant, now more usually blackcurrant and
redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribes r ...
, two berries unrelated to grapes, are derived from this use. Some other fruits of similar appearance are also so named, for example, Australian currant, native currant, Indian currant. A ''sultana'' was originally a raisin made from Sultana grapes of Turkish origin (known as Thompson Seedless in the United States), but the word is now applied to raisins made from either white grapes or red grapes that are bleached to resemble the traditional sultana.


Juice

Grape juice is obtained from crushing and blending grapes into a liquid. The juice is often sold in stores or
fermented Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food p ...
and made into
wine Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...
, brandy, or
vinegar Vinegar is an aqueous solution of acetic acid and trace compounds that may include flavorings. Vinegar typically contains 5–8% acetic acid by volume. Usually, the acetic acid is produced by a double fermentation, converting simple sugars to eth ...
. Grape juice that has been pasteurized, removing any naturally occurring yeast, will not ferment if kept sterile, and thus contains no alcohol. In the
wine Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...
industry, grape juice that contains 7–23% of pulp, skins, stems and seeds is often referred to as "
must Must (from the Latin ''vinum mustum'', "young wine") is freshly crushed fruit juice (usually grape juice) that contains the skins, seeds, and stems of the fruit. The solid portion of the must is called pomace and typically makes up 7–23% of t ...
". In North America, the most common grape juice is purple and made from
Concord grapes The Concord grape is a cultivar derived from the grape species ''Vitis labrusca'' (also known as fox grape) that are used as table grapes, wine grapes and juice grapes. They are often used to make grape Jelly (fruit preserves), jelly, grape juic ...
, while white grape juice is commonly made from Niagara grapes, both of which are varieties of grapes, a different species from European wine grapes. In California, Sultana (known there as Thompson Seedless) grapes are sometimes diverted from the raisin or table market to produce white juice.


Pomace and phytochemicals

Winemaking from red and white grape flesh and skins produces substantial quantities of organic residues, collectively called
pomace Pomace ( ), or marc (; from French ''marc'' ), is the solid remains of grapes, olives, or other fruit after pressing for juice or oil. It contains the skins, pulp, seeds, and stems of the fruit. Grape pomace has traditionally been used to pro ...
(also "marc"), which includes crushed skins, seeds, stems, and leaves generally used as compost. Grape pomace – some 10-30% of the total mass of grapes crushed – contains various phytochemicals, such as unfermented sugars, alcohol,
polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some o ...
s,
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
s, anthocyanins, and numerous other compounds, some of which are harvested and extracted for commercial applications (a process sometimes called "valorization" of the pomace).


Skin

Anthocyanins tend to be the main polyphenolics in purple grapes, whereas
flavan-3-ol Flavan-3-ols (sometimes referred to as flavanols) are a subgroup of flavonoids. They are derivatives of flavans that possess a 2-phenyl-3,4-dihydro-2''H''-chromen-3-ol skeleton. Flavan-3-ols are structurally diverse and include a range of comp ...
s (i.e.
catechin Catechin is a flavan-3-ol, a type of secondary metabolite providing antioxidant roles in plants. It belongs to the subgroup of polyphenols called flavonoids. The name of the catechin chemical family derives from ''catechu'', which is the tannic ...
s) are the more abundant class of polyphenols in white varieties. Total phenolic content is higher in purple varieties due almost entirely to anthocyanin density in purple grape skin compared to absence of anthocyanins in white grape skin. Phenolic content of grape skin varies with
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
, soil composition, climate, geographic origin, and cultivation practices or exposure to diseases, such as fungal infections.
Muscadine ''Vitis rotundifolia'', or muscadine, is a grapevine species native to the southeastern and south-central United States. The growth range extends from Florida to New Jersey coast, and west to eastern Texas and Oklahoma. It has been extensive ...
grapes contain a relatively high phenolic content among dark grapes. In muscadine skins,
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backwards because it can be ...
,
myricetin Myricetin is a member of the flavonoid class of polyphenolic compounds, with antioxidant properties. Common dietary sources include vegetables (including tomatoes), fruits (including oranges), nuts, berries, tea, and red wine. Myricetin is struc ...
,
quercetin Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...
,
kaempferol Kaempferol (3,4′,5,7-tetrahydroxyflavone) is a natural flavonol, a type of flavonoid, found in a variety of plants and plant-derived foods including kale, beans, tea, spinach, and broccoli. Kaempferol is a yellow crystalline solid with a meltin ...
, and trans-resveratrol are major phenolics. The flavonols syringetin, syringetin 3-O-galactoside, laricitrin and laricitrin 3-O-galactoside are also found in purple grape but absent in white grape.


Seeds

Muscadine grape seeds contain about twice the total polyphenol content of skins.
Grape seed oil Grape seed oil (also called grapeseed oil or grape oil) is a vegetable oil derived from the seeds of grapes. A by-product of the winemaking industry, it is typically used for edible applications. Uses Cooking Grape seed oil has a moderatel ...
from crushed seeds is used in
cosmeceutical Cosmeceuticals are cosmetic products with bioactive ingredients purported to have medical benefits. In America, there are no legal requirements to prove that these products live up to their claims. The name is a portmanteau of "cosmetics" and " ...
s and
skincare Skin care is a range of practices that support skin integrity, enhance its appearance, and relieve skin conditions. They can include nutrition, avoidance of excessive sun exposure, and appropriate use of emollients. Practices that enhance appea ...
products. Grape seed oil, including
tocopherol Tocopherols (; TCP) are a class of organic chemical compounds (more precisely, various methylated phenols), many of which have vitamin E activity. Because the vitamin activity was first identified in 1936 from a dietary fertility factor in rat ...
s (
vitamin E Vitamin E is a group of eight fat soluble compounds that include four tocopherols and four tocotrienols. Vitamin E deficiency, which is rare and usually due to an underlying problem with digesting dietary fat rather than from a diet low in vi ...
) and high contents of
phytosterol Phytosterols are phytosteroids, similar to cholesterol, that serve as structural components of biological membranes of plants. They encompass plant sterols and stanols. More than 250 sterols and related compounds have been identified. Free phy ...
s and
polyunsaturated fatty acid Polyunsaturated fatty acids (PUFAs) are fatty acids that contain more than one double bond in their backbone. This class includes many important compounds, such as essential fatty acids and those that give drying oils their characteristic proper ...
s such as
linoleic acid Linoleic acid (LA) is an organic compound with the formula COOH(CH2)7CH=CHCH2CH=CH(CH2)4CH3. Both alkene groups are ''cis''. It is a fatty acid sometimes denoted 18:2 (n-6) or 18:2 ''cis''-9,12. A linoleate is a salt or ester of this acid. ...
,
oleic acid Oleic acid is a fatty acid that occurs naturally in various animal and vegetable fats and oils. It is an odorless, colorless oil, although commercial samples may be yellowish. In chemical terms, oleic acid is classified as a monounsaturated omeg ...
, and
alpha-linolenic acid ''alpha''-Linolenic acid (ALA), also known as α-Linolenic acid (from Greek ''alpha'' meaning "first" and ''linon'' meaning flax), is an ''n''−3, or omega-3, essential fatty acid. ALA is found in many seeds and oils, including flaxseed, wa ...
.


Resveratrol

Resveratrol, a
stilbene Stilbene may refer to one of the two stereoisomers of 1,2-diphenylethene: * (''E'')-Stilbene (''trans'' isomer) * (''Z'')-Stilbene (''cis'' isomer) See also * Stilbenoid Stilbenoids are hydroxylated derivatives of stilbene. They have a C6– ...
compound, is found in widely varying amounts among grape varieties, primarily in their skins and seeds.
Muscadine ''Vitis rotundifolia'', or muscadine, is a grapevine species native to the southeastern and south-central United States. The growth range extends from Florida to New Jersey coast, and west to eastern Texas and Oklahoma. It has been extensive ...
grapes have about one hundred times higher concentration of stilbenes than pulp. Fresh grape skin contains about 50 to 100 micrograms of resveratrol per gram.


Health claims


French paradox

Comparing diets among Western countries, researchers have discovered that although French people tend to eat higher levels of animal fat, the incidence of heart disease remains low in France. This phenomenon has been termed the French paradox, and is thought to occur from protective benefits of regularly consuming red wine, among other dietary practices. Alcohol consumption in moderation may be cardioprotective by its minor anticoagulant effect and
vasodilation Vasodilation is the widening of blood vessels. It results from relaxation of smooth muscle cells within the vessel walls, in particular in the large veins, large arteries, and smaller arterioles. The process is the opposite of vasoconstrictio ...
. Although adoption of wine consumption is generally not recommended by health authorities, some research indicates moderate consumption, such as one glass of red wine a day for women and two for men, may confer health benefits. Alcohol itself may have protective effects on the cardiovascular system.


Grape and raisin toxicity in dogs

The consumption of grapes and raisins presents a potential health threat to dogs. Their toxicity to dogs can cause the animal to develop
acute kidney failure Acute kidney injury (AKI), previously called acute renal failure (ARF), is a sudden decrease in kidney function that develops within 7 days, as shown by an increase in serum creatinine or a decrease in urine output, or both. Causes of AKI are c ...
(the sudden development of kidney failure) with
anuria Anuria is nonpassage of urine, in practice is defined as passage of less than 100Harrison's Principles of Internal Medicine, 19E PAGE 292 milliliters of urine in a day. Anuria is often caused by failure in the function of kidneys. It may also occu ...
(a lack of urine production) and may be fatal.


In religion

Christians have traditionally used wine during worship services as a means of remembering the blood of Jesus Christ which was shed for the remission of sins. Christians who oppose the partaking of alcoholic beverages sometimes use grape juice as the "cup" or "wine" in the
Lord's Supper The Eucharist (; from Greek , , ), also known as Holy Communion and the Lord's Supper, is a Christian rite that is considered a sacrament in most churches, and as an ordinance in others. According to the New Testament, the rite was instit ...
. The
Catholic Church The Catholic Church, also known as the Roman Catholic Church, is the largest Christian church, with 1.3 billion baptized Catholics worldwide . It is among the world's oldest and largest international institutions, and has played a ...
continues to use wine in the celebration of the
Eucharist The Eucharist (; from Greek , , ), also known as Holy Communion and the Lord's Supper, is a Christian rite that is considered a sacrament in most churches, and as an ordinance in others. According to the New Testament, the rite was instit ...
because it is part of the tradition passed down through the ages starting with Jesus Christ at the
Last Supper Image:The Last Supper - Leonardo Da Vinci - High Resolution 32x16.jpg, 400px, alt=''The Last Supper'' by Leonardo da Vinci - Clickable Image, Depictions of the Last Supper in Christian art have been undertaken by artistic masters for centuries, ...
, where Catholics believe the consecrated bread and wine become the
body Body may refer to: In science * Physical body, an object in physics that represents a large amount, has mass or takes up space * Body (biology), the physical material of an organism * Body plan, the physical features shared by a group of anima ...
and blood of Jesus Christ, a dogma known as
transubstantiation Transubstantiation (Latin: ''transubstantiatio''; Greek: μετουσίωσις '' metousiosis'') is, according to the teaching of the Catholic Church, "the change of the whole substance of bread into the substance of the Body of Christ and of ...
. Wine is used (not grape juice) both due to its strong Scriptural roots, and also to follow the tradition set by the early Christian Church. The Code of Canon Law of the Catholic Church (1983), Canon 924 says that the wine used must be natural, made from grapes of the vine, and not corrupt.


Gallery

File:GrapesBuds.JPG, Flower buds File:GrapesFlowers.JPG, Flowers File:TenderGrapes.JPG, Immature fruit File:Grapes Angoor.JPG, Grapes in Iran File:Grapes.jpg, Wine grapes File:Cyprusgrapefarm.jpg, Vineyard in the
Troodos Mountains Troodos (sometimes spelled Troödos; el, Τρόοδος ; tr, Trodos Dağları) is the largest mountain range in Cyprus, located in roughly the center of the island. Its highest peak is Mount Olympus ( el, Όλυμπος), also known as Chion ...
File:Seedless grapes of Kallidaikurichi.jpg, seedless grapes File:Grapes in the Philippines 1.jpg, Grapes in the La Union, Philippines


See also

*
Annual growth cycle of grapevines The annual growth cycle of grapevines is the process that takes place in the vineyard each year, beginning with bud break in the spring and culminating in leaf fall in autumn followed by winter dormancy. From a winemaking perspective, each step ...
* Drakshasava, a traditional Ayurvedic tonic made from grapes *
Grape syrup Grape syrup is a condiment made with concentrated grape juice. It is thick and sweet because of its high ratio of sugar to water. Grape syrup is made by boiling grapes, removing their skins, squeezing them through a sieve to extract the juice, ...
*
List of grape dishes This is a list of notable grape dishes and foods that are prepared using grapes as a primary ingredient. Raisin dishes and foods are also included in this article. Grape dishes and foods A grape is a fruit, botanically a berry, of the decid ...
*
List of grape varieties This list of grape varieties includes cultivated grapes, whether used for wine, or eating as a table grape, fresh or dried (raisin, currant, sultana). For a complete list of all grape species including those unimportant to agriculture, see Vit ...
* ''
Menispermum canadense ''Menispermum canadense'', the Canadian moonseed, common moonseed, or yellow parilla, is a flowering plant in the family Menispermaceae, native to eastern North America, from southern Canada south to northern Florida, and from the Atlantic coas ...
'', a poisonous species resembling wild grapes *
Propagation of grapevines The propagation of grapevines is an important consideration in commercial viticulture and winemaking. Grapevines, most of which belong to the ''Vitis vinifera'' family, produce one crop of fruit each growing season with a limited life span for ind ...
* The Fox and the Grapes


References


Further reading

* Creasy, G. L. and L. L. Creasy (2009). ''Grapes'' (Crop Production Science in Horticulture). CABI. .


External links

* * {{Authority control Berries Crops originating from Europe Edible fruits Plants in the Bible Flora of Palestine (region) Flora of Israel