Graft-chimera
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In horticulture, a graft-chimaera may arise in
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
at the point of contact between rootstock and scion and will have properties intermediate between those of its "parents". A graft-chimaera is not a true hybrid but a mixture of cells, each with the
genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
of one of its "parents": it is a
chimaera Chimaeras are cartilaginous fish in the order Chimaeriformes , known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively. At ...
. Hence, the once widely used term "graft-hybrid" is not descriptive; it is now frowned upon. Propagation is by cloning only. In practice graft-chimaeras are not noted for their stability and may easily revert to one of the "parents".


Nomenclature

Article 21 of the '' ICNCP'' stipulates that a graft-chimaera can be indicated either by * a formula: the names of both "parents", in alphabetical order, joined by the
plus sign The plus and minus signs, and , are mathematical symbols used to represent the notions of positive and negative, respectively. In addition, represents the operation of addition, which results in a sum, while represents subtraction, result ...
"+": : ''Crataegus'' + ''Mespilus'' * a name: ** if the "parents" belong to different
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
a name may be formed by joining part of one generic name to the whole of the other generic name. This name must not be identical to a generic name published under the ''
International Code of Nomenclature for algae, fungi, and plants The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all those "trad ...
'' (''ICN''). For example + ''Crataegomespilus'' is the name for the graft-chimaera which may also be indicated by the formula ''Crataegus'' + ''Mespilus''. This name is clearly different from ×''Crataemespilus'', the name under the ''ICN'' for the true hybrid between ''Crataegus'' and ''Mespilus'', which can also be designated by the formula ''Crataegus'' × ''Mespilus''. ** if both "parents" belong to the same genus the graft-chimaera may be given a cultivar name. For example ''Syringa'' 'Correlata' is a graft-chimaera involving ''Syringa vulgaris'' (common lilac) and ''
Syringa × chinensis ''Syringa'' × ''chinensis'', the Chinese lilac or Rouen lilac, is a hybrid species of flowering plant in the family Oleaceae. It was supposedly first noticed growing in Rouen, France in 1777. In spite of its specific and common names, it most p ...
'' (Rouen lilac, which is itself a hybrid between ''S. vulgaris'' and ''S. persica''). No plus sign is used, because both "parents" belong to the genus ''Syringa''. A graft-chimaera cannot have a species name, because it is simultaneously two species. Although +''Laburnocytisus'' 'Adamii', for example, is sometimes seen written as if it were a species (+''Laburnocytisus adamii''), this is incorrect.


In Darwin's works

Charles Darwi
"The Variation of Animals and Plants Under Domestication"
, 1868 г.:
I will therefore give all the facts which I have been able to collect on the formation of hybrids between distinct species or varieties, without the intervention of the sexual organs. For if, as I am now convinced, this is possible, it is a most important fact, which will sooner or later change the views held by physiologists with respect to sexual reproduction. A sufficient body of facts will afterwards be adduced, showing that the segregation or separation of the characters of the two parent-forms by bud-variation, as in the case of Cytisus adami, is not an unusual though a striking phenomenon. We shall further see that a whole bud may thus revert, or only half, or some smaller segment.


See also

* Bizzaria – a very popular graft chimaera in ''Citrus'' * *
Grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...


References

{{Reflist Botanical nomenclature