Goniothalamus Ridleyi
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''Goniothalamus ridleyi'' is a species of
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
in the family
Annonaceae The Annonaceae are a family of flowering plants consisting of trees, shrubs, or rarely lianas commonly known as the custard apple family or soursop family. With 108 accepted genera and about 2400 known species, it is the largest family in the Ma ...
. It is native to
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and ea ...
,
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
, Sumatra and
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
.
George King George King may refer to: Politics * George King (Australian politician) (1814–1894), New South Wales and Queensland politician * George King, 3rd Earl of Kingston (1771–1839), Irish nobleman and MP for County Roscommon * George Clift King (18 ...
, who first formally described the species, named it after the English botanist
Henry Nicholas Ridley Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees i ...
who collected the specimen King examined.


Description

It is a tree reaching 23 meters in height. Its papery to membranous, elliptical leaves are 14-29 by 3.5-11.4 centimeter, have short tapering tips and bases that come to a point where they meet the petioles. The dull upper surfaces of the leaves are hairless except the midribs which have fine hairs. The shiny lower surfaces are hairless except the midribs which have fine hairs. The leaves have about 6-19 pairs of secondary veins that emanate from their midribs that curve near the leaf margin. Its petioles are 6-16 by 1-3 millimeters and either hairless or sparse with fine hairs. Its flowers occur in crowded clusters on long
pedicels In botany, a pedicel is a stem that attaches a single flower to the inflorescence. Such inflorescences are described as ''pedicellate''. Description Pedicel refers to a structure connecting a single flower to its inflorescence. In the absenc ...
that extend from warty outgrowths, or
tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...
, that grow from the base of the trunk close to soil level. The tubercles are woody and covered in fine hairs. The sparsely hairy pedicels are 1.5-11 centimeters by 1.2-1.9 millimeters and have 3-10 small bracts. Its purple to brown, leathery, oval to triangular
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s are 9–1.35 by 6-9 millimeters, sparsely covered in fine hairs, with bluntly pointed tips. The sepals are fused at their base. Its flowers have 6 petals in two rows of 3. The cream-colored to pale brown, leathery, oblong to oval outer petals are 1.2-5 by 0.9-2 centimeters, have broad thick
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...
s, come to shallow point at their tips. The out petals are covered in wooly hairs on their inner surface and sparse to dense fine hairs on their outer surface. The cream-colored to pale brown, leathery, teardrop-shaped inner leaves are 1.1-1.7 by 0.7-0.95 centimeters, taper to a short slender point at their tips, and have claws that are 1.5-2.4 millimeters wide. The inner leaves densely hairy on their outer surface and velvety on their inner surface. Its flowers have 120-300 stamen that are 1.5-3.4 by 0.2-0.4 millimeters. Its flowers have 34-90 carpels with hairless 0.9–2.3 by 0.2–0.5 millimeter
ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...
. The stigma and style are 1-2 by 0.1-0.3 millimeters long and hairless or sparse with fine hairs. Its fruit are born on pedicels that are 3-13 centimeters by 2-5 millimeters. Its smooth, matt, hairless, round to oval, fruit are 1.6-3.5 by 1.3-2.9 centimeters and tapered. The fruit vary in color including yellow, pink, purple and red to brown. Each fruit is attached to the pedicel by a 2-15 by 1.5-3 millimeter stipe. The fruit have 1-3 flattened, elliptical seeds that are 1.5-2.5 by 1-1.8 centimeters. The brown seeds are smooth and sparse tufts of white or gold-colored hairs.


Reproductive biology

The pollen of ''G. ridleyi'' is shed as permanent tetrads.


Habitat and distribution

It has been observed growing in swamp forests, lowland forests and montane forests, on hillside or by streams, at elevations of 50 to 1000 meters.


References

{{Taxonbar, from=Q15359223
ridleyi Henry Nicholas Ridley CMG (1911), MA (Oxon), FRS, FLS, F.R.H.S. (10 December 1855 – 24 October 1956) was an English botanist, geologist and naturalist who lived much of his life in Singapore. He was instrumental in promoting rubber trees ...
Flora of Borneo Flora of Peninsular Malaysia Flora of Sumatra Flora of Thailand Plants described in 1892 Taxa named by George King (botanist)