Goniasteridae - Fromia Monilis
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Goniasteridae (the biscuit stars) constitute the largest
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
sea star Starfish or sea stars are Star polygon, star-shaped echinoderms belonging to the class (biology), class Asteroidea (). Common usage frequently finds these names being also applied to brittle star, ophiuroids, which are correctly referred to ...
s, included in the order
Valvatida The Valvatida are an order of starfish in the class Asteroidea, which contains 695 species in 172 genera in 17 families. Description The order encompasses both tiny species, which are only a few millimetres in diameter, like those in the genu ...
. They are mostly deep-dwelling species, but the family also include several colorful shallow tropical species.


Description

Goniasteridae are usually middle-sized sea stars with a characteristic double range of marginal plates bordering the disk and arms. Most of them have five arms, often short and triangular, around a broad central disc; many species are pentagonal or subpentagonal, covered densely with granular, seed-like protuberances, hence the name of the family "seed-star" (''gonium+aster''). The aboral face is often covered with tiny spines looking like paxillae.
Pedicellaria A pedicellaria (plural: pedicellariae) is a small wrench- or claw-shaped appendage with movable jaws, called valves, commonly found on echinoderms (phylum Echinodermata), particularly in sea stars (class Asteroidea) and sea urchins (class Echinoi ...
e are often valvate, and the gonads are located at the interradius. Main identification keys for this group include the presence of paxillae, granules, teeth, spines, or the shape and dimensions of marginal plate.


Location and habitat

They occur predominantly on deep-water
continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
habitats (but a part of them inhabit shallow waters) in all the world's oceans, being the most diverse in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region.


List of genera

About 260 extant species within 70 genera are currently known, which make this family the most diverse of all the sea stars, even if half of the genera are monospecific. Species belonging to the '' Ferdininae'' subfamily have been imported from
Ophidiasteridae The Ophidiasteridae (Greek ''ophidia'', Οφιδια, "of snakes", diminutive form) are a family of sea stars with about 30 genera. Occurring both in the Indo-Pacific and Atlantic Oceans, ophidiasterids are greatest in diversity in the Indo-Pacif ...
thanks to a large revision of these two families in 2017 According to
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
, this family includes the following genera: Image:Akelbaster caledoniae.JPG, '' Akelbaster novaecaledoniae'' Image:Anthenoides peircei - NOAA Photo Library.jpg, '' Anthenoides piercei'' Image:Apollonaster kelleyi.jpg, '' Apollonaster kelleyi'' Image:Astroceramus eldredgei.jpg, '' Astroceramus eldredgei'' Image:Calliaster baccatus P3164673.JPG, ''
Calliaster baccatus ''Calliaster baccatus'', the cobbled starfish, is a species of starfish in the family Goniasteridae.Clark A. & Courtman-Stock J. 1976. ''The Echinoderms of South Africa'' British Museum (Natural History) London Description The cobbled starfi ...
'' Image:Ceramaster granularis-25mm.jpg, '' Ceramaster granularis'' Image:Chitonaster johannae.jpg, '' Chitonaster johannae'' Image:Circeaster sandrae.JPG, '' Circeaster sandrae'' Image:Cladaster analogus.JPG, '' Cladaster analogus'' Image:Eknomiaster beccae.JPG, '' Eknomiaster beccae'' Image:Eratosaster jenae.JPG, '' Eratosaster jennae'' Image:Evoplosoma.jpg, '' Evoplosoma sp.'' Image:Eosaster nadiae.jpg, '' Eosaster nadiae'' Image:Ferdina flavescens.jpg, '' Ferdina flavescens'' Image:Kanakaster solidus.jpg, '' Kanakaster solidus'' Image:Neoferdina cumingi.jpg, ''
Neoferdina cumingi ''Neoferdina cumingi'', also known as Cuming's sea star, is a species of starfish in the Family (biology), family Goniasteridae. It is native to the tropical Indo-Pacific region. Description This starfish has a wide variety of colour forms but t ...
'' Image:Paraferdina plakos.jpg, '' Paraferdina plakos'' Image:Fromia monilis (Seastar).jpg, ''
Fromia monilis ''Fromia monilis'', common name necklace starfish or tiled starfish, is a species of starfish belonging to the family Goniasteridae. Description ''Fromia monilis'' can reach a diameter of about . Tips of the arms and the disc center of this sta ...
'' Image:Gilbertaster caribaea.jpg, '' Gilbertaster caribaea'' Image:Glyphodiscus magnificus.jpg, '' Glyphodiscus magnificus'' Image:Goniaster tessellatus São Tomé.jpg, ''
Goniaster tessellatus ''Goniaster'' is a genus of echinoderms belonging to the family Goniasteridae Goniasteridae (the biscuit stars) constitute the largest family of sea stars, included in the order Valvatida. They are mostly deep-dwelling species, but the family a ...
'' Image:Olympic Coast National Marine Sanctuary 2010 Hippasteria spinosa.jpg, '' Hippasteria spinosa'' Image:Iconaster longimanus.jpg, '' Iconaster longimanus'' Image:Mariaster giganteus 01.jpg, `''Mariaster giganteus'' Image:Mediaster aequalis 25March2010 14.jpg, '' Mediaster aequalis'' Image:Milteliphaster woodmasoni.jpg, '' Milteliphaster woodmasoni'' Image:Nectria macrobrachia Large plated seastar P1021082.JPG, '' Nectria macrobrachia'' Image:Nymphaster arenatus.png, '' Nymphaster arenatus'' Image:Peltaster placenta.jpg, ''
Peltaster placenta ''Peltaster'' is a genus of echinoderms belonging to the family Goniasteridae. The genus has almost cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of ...
'' Image:Pawsonaster parvus.jpg, '' Pawsonaster parvus'' Image:Pentagonaster duebeni 1.jpg, ''
Pentagonaster duebeni ''Pentagonaster duebeni'' is a species of starfish within the family Goniasteridae. Distribution & habitat ''Pentagonaster duebeni'' is distributed off the Australian coasts from Shark Bay to Tasmania and southern Queensland. Its preferred hab ...
'' Image:Pillsburiaster calvus.JPG, '' Pillsburiaster calvus'' Image:Plinthaster dentatus.jpg, '' Plinthaster dentatus'' Image:Pseudoceramaster regularis.jpg, '' Pseudoceramaster regularis'' Image:Rosaster.jpg, '' Rosaster sp.'' Image:Ryukuaster onnae.JPG, '' Ryukuaster onnae'' Image:Sphaeriodiscus mirabilis.JPG, ''
Sphaeriodiscus mirabilis ''Sphaeriodiscus'' is a genus of echinoderms belonging to the family Goniasteridae. The genus has almost cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or mos ...
'' Image:Galloping sea star (Stellaster equestris).jpg, '' Stellaster childreni'' Image:Stellasteropsis fouadi (MNHN-IE-2014-49) 01.jpg, ''
Stellasteropsis fouadi ''Stellasteropsis'' is a genus of echinoderms belonging to the family Goniasteridae. The species of this genus are found in Indian Ocean and Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indom ...
'' Image:Sthenaster emmae.JPG, ''
Sthenaster emmae ''Sthenaster emmae'' is a species of deep sea corallivorous (preys on deep-sea corals) sea star. It is the only known species in the genus ''Sthenaster''. This species in particular is found in the tropical Atlantic. Description ''Sthenaste ...
'' Image:Tosia australis P1222858.JPG, '' Tosia australis''


Extinct genera

Lists of genera containing extinct species according to fossilworks.


References

{{Taxonbar, from=Q2076886 Echinoderm families Extant Jurassic first appearances