Gongbusaurus
   HOME

TheInfoList



OR:

''Gongbusaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
n, perhaps
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous world ...
,
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived between about 160 and 157 million years ago, in the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
. A small
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, it is very poorly known. Two species have been assigned to it, but as the original name is based on teeth, there is no concrete evidence to connect the two species. Its fossils have been found in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.


Description

''Gongbusaurus'', by extrapolation from the remains of possible species "G." ''wucaiwanensis'' and other basal ornithopods, was a herbivorous
bipedal Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
animal around long. The
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
of "G." ''wucaiwanensis'' is 19.5 centimetres long. It would have been a strong runner.


Classification

Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at the ...
, Zhou Shiwu, and Zhang Zicheng, who originally described the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''G. shiyii'', thought it was most similar to ''
Fabrosaurus ''Fabrosaurus'' ( ) is an extinct genus of ornithischian dinosaur that lived during the Early Jurassic during the Hettangian to Sinemurian stages 199 - 189 mya.Ginsburg, L., (1964), "Decouverte d’un Scelidosaurien (Dinosaure ornithischien) ...
'' and assigned it to the nebulous
Fabrosauridae Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
. Upon description of the second species "G." ''wucaiwanensis'' several years later, Dong elected to assign it to Hypsilophodontidae, an equally nebulous (
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
) family of somewhat more
derived Derive may refer to: * Derive (computer algebra system), a commercial system made by Texas Instruments * ''Dérive'' (magazine), an Austrian science magazine on urbanism *Dérive, a psychogeographical concept See also * *Derivation (disambiguatio ...
small bipedal
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
ns, while at about the same time,
David B. Weishampel Professor David Bruce Weishampel (born November 16, 1952) is an American palaeontologist in the Center for Functional Anatomy and Evolution at Johns Hopkins University School of Medicine. Weishampel received his Ph.D. in Geology from the Universi ...
and Larry Witmer found ''Gongbusaurus'' to be an indeterminate basal ornithischian. The most recent reviews also found the genus to be a
dubious Doubt is a mental state in which the mind remains suspended between two or more contradictory propositions, unable to be certain of any of them. Doubt on an emotional level is indecision between belief and disbelief. It may involve uncertainty, ...
ornithischian, and recommended renaming the better-known second species.
Peter Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosaur ...
has noted that the teeth on which ''Gongbusaurus'' is based resemble those of ''
Sarcolestes ''Sarcolestes'' (meaning "flesh robber") is an extinct genus of ankylosaurian ornithischian dinosaur from the Oxford Clay of England. The current type and only species is ''S. leedsi'', and the holotype is a single partial left mandible. The ...
'' and '' Gastonia'', so the genus may actually be an
ankylosauria Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
n.


History

Dong and his coauthors established ''Gongbusaurus'' on two small teeth,
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
IVPP V9069: one, V9069-1, from the beak (
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
), and the other, V9069-2, from the cheek (
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
). These remains came from the Oxfordian-age
Upper Shaximiao Formation The Shaximiao Formation () is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the ...
in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
, China and were found by a unit of the Chinese aerial survey behind an elementary school in the village of Huangtong, part of the Duxin commune, in Rongxian or Rong County. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Gongbusaurus shiyii'' was named and described by Dong, Zhou, and Zhang in 1983. The generic name is derived from the imperial ministry, ''bu'', of public works, ''gong'', in honour of the great poet
Du Fu Du Fu (; 712–770) was a Tang dynasty poet and politician. Along with his elder contemporary and friend Li Bai (Li Po), he is frequently called the greatest of the Chinese poets.Ebrey, 103. His greatest ambition was to serve his country as ...
who in Sichuan worked for that ministry after 760, in the rank of ''shi yi'', "junior consultant", which explains the specific name. As ''shi yi'' also can mean "register of lost objects" the specific name is at the same time a pun on the fact that the teeth were later separately found among the assorted bones collected during the dig. Dong added a second species, "G." ''wucaiwanensis'', in 1989 for a fragmentary skeleton (holotype IVPP 8302) including a partial lower jaw, three tail
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, and a partial forelimb, and added another specimen (
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Of ...
IVPP 8303) consisting of two hip vertebrae, eight tail vertebrae, and two complete hind limbs. A partial foot (IVPP 8304) and four separate dorsal vertebrae and a caudal vertebra were referred. These remains came from the roughly contemporaneous
Shishugou Formation The Shishugou Formation () is a geological formation in Xinjiang, China. Its strata date back to the Late Jurassic period. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Din ...
of Wucaiwan,
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
. Tooth species are not well regarded in dinosaur paleontology, because dinosaur teeth are generally not distinctive enough to hold a name. Therefore, it is unsurprising that other paleontologists have suggested removing "G." ''wucaiwanensis''. A possible replacement name, " Eugongbusaurus", has accidentally gotten to the public, but remains
informal Formal, formality, informal or informality imply the complying with, or not complying with, some set of requirements (forms, in Ancient Greek). They may refer to: Dress code and events * Formal wear, attire for formal events * Semi-formal attire ...
.


References

{{Taxonbar, from=Q1656761 Ornithischian genera Late Jurassic dinosaurs of Asia Paleontology in Sichuan Nomina dubia Taxa named by Dong Zhiming Fossil taxa described in 1983