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''Gonepteryx cleopatra'', the Cleopatra or Cleopatra butterfly, is a medium-sized
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
of the family
Pieridae The Pieridae are a large family of butterflies with about 76 genera containing about 1,100 species, mostly from tropical Africa and tropical Asia with some varieties in the more northern regions of North America and Eurasia.DeVries P. J. in Le ...
.


Subspecies

The species ''Gonepteryx cleopatra'' is divided into the following ten
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: *''G. c. cleopatra'' (Linnaeus, 1767)
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
*''G. c. balearica'' Bubacek, 1920
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
*''G. c. petronella'' De Freina, 1977
Ibiza Ibiza (natively and officially in ca, Eivissa, ) is a Spanish island in the Mediterranean Sea off the eastern coast of the Iberian Peninsula. It is from the city of Valencia. It is the third largest of the Balearic Islands, in Spain. Its l ...
*''G. c. italica'' (Gerhardt, 1882)
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Corsica Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...
,
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
*''G. c. dalmatica'' Verity, 1908
Dalmatia Dalmatia (; hr, Dalmacija ; it, Dalmazia; see #Name, names in other languages) is one of the four historical region, historical regions of Croatia, alongside Croatia proper, Slavonia, and Istria. Dalmatia is a narrow belt of the east shore of ...
n coast, western
Balkan The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
*''G. c. citrina'' Sheljuzhko, 1925 – southern
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
*''G. c. insularis'' Verity, 1909
Crete Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the Greek islands, the 88th largest island in the world and the fifth largest island in the Mediterranean Sea, after Sicily, Sardinia, Cyprus, and ...
*''G. c. fiorii'' Turati & Fiori, 1930
Rhodes Rhodes (; el, Ρόδος , translit=Ródos ) is the largest and the historical capital of the Dodecanese islands of Greece. Administratively, the island forms a separate municipality within the Rhodes regional unit, which is part of the So ...
*''G. c. taurica'' (Staudinger, 1881)
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
*''G. c. palmata'' Turati, 1921
Cyrenaica Cyrenaica ( ) or Kyrenaika ( ar, برقة, Barqah, grc-koi, Κυρηναϊκή παρχίαKurēnaïkḗ parkhíā}, after the city of Cyrene), is the eastern region of Libya. Cyrenaica includes all of the eastern part of Libya between ...
,
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to Egypt–Libya bo ...


Appearance

''Gonepteryx cleopatra'' is a medium-sized butterfly with a wingspan of about 50–70 mm (2.0-2.8 in). It is a
sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
species - the female has pale yellow or greenish wings, the male is darker yellow with an orange patch on the forewing. Both sexes have a forewing apical hook and brown dots in the center of each wing, and the underside of wings is light greenish yellow. The greenish color, the shape and the pronounced venation on the hindwings give to these butterflies a good camouflage, making them resemble just leaves.


Ecology

The Cleopatra butterfly inhabits open woodland and scrub. The flight period is from May to August in most parts of its range, except Spain, where it is double brooded and may fly almost all year. The adult hibernates in evergreen trees and shrubs. The caterpillars feed on the
buckthorn ''Rhamnus'' is a genus of about 110 accepted species of shrubs or small trees, commonly known as buckthorns, in the family Rhamnaceae. Its species range from tall (rarely to ) and are native mainly in east Asia and North America, but found thr ...
s ''
Rhamnus alaternus ''Rhamnus alaternus'' is a species of flowering plant in the buckthorn family Rhamnaceae, known by the common names Italian buckthorn or Mediterranean buckthorn. It is a hardy medium-sized evergreen shrub with fragrant flowers. Etymology The s ...
''. When the green caterpillars are prepared to pass to the pupal stage, they tie themselves through a silk belt to the host plant, on the underside of a leaf. The green chrysalis became gradually yellow and red, revealing the colors on the wings of the adult close to flutter.


Distribution

This species is native to the
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
region (
Southern Europe Southern Europe is the southern regions of Europe, region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countrie ...
,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
and
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
).


References

* Tristan Lafranchis
Quand les papillons changent d'habitat
* Bernard d'Abrera, Butterflies of the Holarctic Region, Part I, Hill House, Victoria 1990, S.98 * Tom Tolman/Richard Lewington, Collins Butterfly Guide, HarperCollins, London 1997




Lepinet.fr


External links





{{Taxonbar, from=Q938502 Gonepteryx Butterflies of Africa Butterflies of Asia Butterflies of Europe Butterflies described in 1767 Taxa named by Carl Linnaeus