Golden-tailed Sapphire
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The golden-tailed sapphire (''Chrysuronia oenone'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the family
Trochilidae Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics arou ...
. It is found in
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
, and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
.


Taxonomy and systematics

The golden-tailed sapphire was formerly placed in the genus ''Amazilia''. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2014 found that ''Amazilia'' was
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
. In the revised classification to create
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
genera, the golden-tailed sapphire was moved by most taxonomic systems to ''
Chrysuronia ''Chrysuronia'' is a genus of hummingbirds in the family Trochilidae. Taxonomy The genus ''Chrysuronia'' was introduced in 1850 by the French naturalist Charles Lucien Bonaparte. Bonaparte did not specify a type species but this was designated a ...
''.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 However,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
'' (HBW) retains it in ''Amazilia''. The
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
(IOC) recognizes two subspecies of golden-tailed sapphire, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In the ...
''C. o. oenone'' and ''C. o. josephinae''. The
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
and HBW add a third, ''C. o. alleni'', that the IOC includes within ''josephinae''.


Description

The golden-tailed sapphire is long. Males weigh and females . Both sexes of all subspecies have a slightly curved bill with a black
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
and a black-tipped red
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
. Adult males of the nominate subspecies have a violet-blue head and throat. Their back is bright green that transitions through bronze-green to bright coppery
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
and a copper-bronze tail. Their breast is glittering green, their belly bronze-green, and underneath their tail, their coverts are bronze with whitish edges. Adult females have a blue crown, a green back, and copper-bronze coverts and tail. Their undersides are mostly whitish, with blue-green flecks on the sides of the throat and neck, green on the sides and chest, bronze-green flanks, and underneath their tail, they have bronze coverts. Juvenile males have a dull green crown, a dusky gray throat, and a dull green chest; early adult feathers on the head and throat are a purer, less violet, blue than the adult's. Juvenile females' throat speckles are duller and more bronze-coloured than the adults'.Stiles, F.G. and P. F. D. Boesman (2020). Golden-tailed Sapphire (''Chrysuronia oenone''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.gotsap1.01 retrieved September 12, 2022 Subspecies ''C. o. josephinae'' and ''C. o. alleni'' when treated separately have some differences from the nominate and from each other. Males of the former have a mostly green throat and a green rump that contrasts with the coppery coverts. ''C. o. alleni'' has an entirely green throat and little blue on the cheeks and its rump is coppery.


Distribution and habitat

The nominate subspecies of golden-tailed sapphire is the most widely distributed. It is found in the
Serranía del Perijá The Serranía del Perijá, Cordillera de Perijá or Sierra de Perijá is a mountain range, an extension of the eastern Andean branch ( Cordillera Oriental), in northern South America, between Colombia and Venezuela Venezuela (; ), offici ...
that straddles the Colombia/Venezuela border, further east in western and northern Venezuela, and south through east central Colombia, eastern Ecuador, and extreme western Brazil into extreme northeastern Peru. ''C. o. josephinae''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
is found in most of the rest of eastern Peru. ''C. o. alleni'' is found in northern Bolivia. The golden-tailed sapphire inhabits semi-open landscapes like the edges and gaps in humid forest, mature
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
,
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, shady cacao and coffee plantations, and gardens. In Colombia and Venezuela it ranges in elevation from sea level to and in Ecuador between .


Behavior


Movement

The golden-tailed sapphire apparently moves locally according to the availability of flowering plants but details are lacking.


Feeding

The golden-tailed sapphire forages for nectar at a variety of flowering shrubs (e.g. ''
Aphelandra ''Aphelandra'' is a genus of about 170 species of flowering plants in the family Acanthaceae, native to tropical regions of the Americas. They are evergreen shrubs growing to tall, with opposite, simple leaves long, often with white veins. The ...
'') and trees (e.g. ''
Erythrina ''Erythrina'' is a genus of plants in the pea family, Fabaceae. It contains about 130 species, which are distributed in tropical and subtropical regions worldwide. They are trees, with the larger species growing up to in height. The generic na ...
'' and ''
Inga ''Inga'' is a genus of small tropical, tough-leaved, nitrogen-fixing treesElkan, Daniel. "Slash-and-burn farming has become a major threat to the world's rainforest" ''The Guardian'' 21 April 2004 and shrubs, subfamily Mimosoideae. ''Inga''s l ...
''). Males tend to forage in the canopy, sometimes gathering in "quarrelsome assemblages", while females forage more by
trap-lining In ethology and behavioral ecology, trap-lining or traplining is a feeding strategy in which an individual visits food sources on a regular, repeatable sequence, much as trappers check their lines of traps. Traplining is usually seen in species ...
in the forest, visiting a circuit of flowering plants. In addition to nectar, it feeds on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist *Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Haw ...
from a perch or by gleaning from foliage.


Breeding

The golden-tailed sapphire's breeding season varies geographically. It spans at least June to November in eastern Colombia but is January through April in western Brazil. Males usually gather in groups of five to 10 at
leks A lek is an aggregation of male animals gathered to engage in competitive displays and courtship rituals, known as lekking, to entice visiting females which are surveying prospective partners with which to mate. A lek can also indicate an avail ...
to display to females but sometimes display alone. The nest is cup-shaped and has green moss on the outside for camouflage. The female incubates the clutch of two eggs for about 14 days and fledging occurs about 28 days after hatch.


Vocalization

The golden-tailed sapphire's song is variable, described as "a repeated rhythmic phrase, often starting with a burry note, followed by several very squeaky or scratchy notes, and ending with repeated chips." It also makes calls described as "a drawn-out metallic trill and shorter chipping notes."


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the golden-tailed sapphire as being of Least Concern. It has a very large range, and though its population size is unknown it is believed to be stable. No immediate threats have been identified. It is considered locally common to common in Colombia and Venezuela but relatively rare in the Amazonian lowlands further south.


References

{{Taxonbar, from=Q775398
golden-tailed sapphire The golden-tailed sapphire (''Chrysuronia oenone'') is a species of hummingbird in the family Trochilidae. It is found in Bolivia, Brazil, Colombia, Ecuador, Peru, and Venezuela. Taxonomy and systematics The golden-tailed sapphire was formerl ...
Hummingbird species of South America Birds of Colombia Birds of the Venezuelan Andes Birds of the Venezuelan Coastal Range Birds of the Ecuadorian Amazon Birds of the Peruvian Amazon Birds of the Bolivian Amazon
golden-tailed sapphire The golden-tailed sapphire (''Chrysuronia oenone'') is a species of hummingbird in the family Trochilidae. It is found in Bolivia, Brazil, Colombia, Ecuador, Peru, and Venezuela. Taxonomy and systematics The golden-tailed sapphire was formerl ...
Taxa named by René Lesson Taxonomy articles created by Polbot Taxobox binomials not recognized by IUCN