Goldbachia
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''Goldbachia'' is a genus of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s belonging to the family Brassicaceae.


General description

Goldbachia are annual, herbaceous plants. They lack trichomes (hairs) and have branched stems. The basal leaves are
petiolate Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, a ...
(without stalk), entire, and dentate (with toothed margins). Some species are rosulate (in a rosette), while others are not. The cauline leaves are
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, auriculate (rarely they are not) or amplexicaul (clasping the stem) at the base, and entire, with the margins either repand (a slightly undulating margin) or dentate. The
cotyledons A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...
(primary leaves) are incumbent. The
racemes A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
are ebracteate (having no bracts) and elongated when in fruit. Fruiting pedicels are filiform (thread-like), recurved or reflexed, and articulate at the base or both ends. The
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
are ovate or oblong shaped, suberect, base of lateral pair, and either not saccate (pouch shaped) or sub-saccate. The petals are white, pink, or purplish and are longer than the sepals. The blade is spatulate (spoon shaped) with an obtuse apex. The claw is different than the sepals. It has 6 stamens, which are slightly tetradynamous (four of which are longer than the others). The median filament pairs are flattened at the base, with the lateral pair filiform. The
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
are ovate and obtuse at the apex. It has 4
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
glands, which are minute; the median glands are free or confluent with laterals. The lateral glands are semi-annular and intrastaminal (inside the stamens). The ovules are 1-3 per ovary, and are subapical. The fruit (seed capsule) is an indehiscent nutlet-like silicle or silicula, often breaking into 1-seeded units, and are oblong, ovoid, or ellipsoid. They are 1-loculed or transversely 2- or 3-loculed, terete (circular in cross-section) or 4-angled,
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, woody, often prominently veined, glabrous or papillate (paper-like), smooth or torulose (a cylindrical or ellipsoid body; swollen and constricted at intervals). It is sometimes verrucose-reticulate; replum (a framework-like placenta) rounded; septum absent; style absent, obsolete, or distinct, thickened and sub-conical; stigma capitate, entire. The
seeds A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm pl ...
are uniseriate (arranged in a single row), wingless, oblong and plump. The seed coat is smooth, and not mucilaginous (having a viscous or gelatinous consistency) when damp.


Taxonomy

The genus name of ''Goldbachia'' is in honour of Carl Ludwig Goldbach (1793–1824), German-born Russian professor of botany in
Moscow Moscow ( , US chiefly ; rus, links=no, Москва, r=Moskva, p=mɐskˈva, a=Москва.ogg) is the capital and largest city of Russia. The city stands on the Moskva River in Central Russia, with a population estimated at 13.0 million ...
, and it was first described and published by Augustin Pyramus de Candolle in
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
's book Syst. Nat. Vol.2 on page 576 in 1821.


Known species

Includes 7 accepted Species, according to Kew; The genus was verified by
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
and the
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
on 31 July 2018.


Range and habitat

Its native range is eastern Europe,
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
, to Mongolia and Pakistan. It is found in the countries of Afghanistan, China (including the regions of Xinjiang, Inner Mongolia and Qinghai), Iran, Iraq, Kazakhstan, Kyrgyzstan, Lebanon-Syria, Mongolia, Pakistan, Saudi Arabia, Tajikistan, Tibet, Turkey, Turkmenistan,
Tuva Tuva (; russian: Тува́) or Tyva ( tyv, Тыва), officially the Republic of Tuva (russian: Респу́блика Тыва́, r=Respublika Tyva, p=rʲɪˈspublʲɪkə tɨˈva; tyv, Тыва Республика, translit=Tyva Respublika ...
(part of Russia), Uzbekistan, and western Siberia. ''Goldbachia laevigata'', ''Goldbachia pendula'' and ''Goldbachia ikonnikovii'' are found in China.


References

{{Taxonbar, from=Q3081963 Brassicaceae Brassicaceae genera Plants described in 1821 Flora of the Indian subcontinent Flora of Western Asia Flora of the Caucasus Flora of Mongolia