Gojirasaurus
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''Gojirasaurs'' (meaning "
Godzilla is a fictional monster, or '' kaiju'', originating from a series of Japanese films. The character first appeared in the 1954 film ''Godzilla'' and became a worldwide pop culture icon, appearing in various media, including 32 films produc ...
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
")K. Carpenter, 1997, "A giant coelophysoid (Ceratosauria) theropod from the Upper Triassic of New Mexico, USA", ''Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen'' 205 (#2): 189-208 is genus of
coelophysoid Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
theropod dinosaur named after the giant monster movie character Gojira (the Hepburn romanized
Japanese Japanese may refer to: * Something from or related to Japan, an island country in East Asia * Japanese language, spoken mainly in Japan * Japanese people, the ethnic group that identifies with Japan through ancestry or culture ** Japanese diaspor ...
name for the monster
Godzilla is a fictional monster, or '' kaiju'', originating from a series of Japanese films. The character first appeared in the 1954 film ''Godzilla'' and became a worldwide pop culture icon, appearing in various media, including 32 films produc ...
).


Etymology

The composite term ''Gojirasaurus'' is derived from the name of the giant Japanese movie monster "Gojira" (
Godzilla is a fictional monster, or '' kaiju'', originating from a series of Japanese films. The character first appeared in the 1954 film ''Godzilla'' and became a worldwide pop culture icon, appearing in various media, including 32 films produc ...
) and the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word "" () meaning "lizard"; thus, "Godzilla lizard". In addition, a theropod dinosaur with the name ''Godzillasaurus'' exists in the Heisei era of ''Godzilla'' films, and in that continuity is explained to be the non mutated form of Godzilla. "Gojira" was selected as a reference to the great size of this theropod, which exceeded that of its Triassic counterparts. The specific name ''quayi'', is a reference to Quay County,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
, where the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen was discovered. ''Gojirasaurus'' was described and named by
Kenneth Carpenter Kenneth Carpenter (born September 21, 1949, in Tokyo, Japan) is a paleontologist. He is the former director of the USU Eastern Prehistoric Museum and author or co-author of books on dinosaurs and Mesozoic life. His main research interests ar ...
in
1997 File:1997 Events Collage.png, From left, clockwise: The movie set of ''Titanic'', the highest-grossing movie in history at the time; ''Harry Potter and the Philosopher's Stone'', is published; Comet Hale-Bopp passes by Earth and becomes one of t ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Gojirasaurus quayi''.


Description

''Gojirasaurus'' is one of the largest meat-eating dinosaurs known from the
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
Period, almost the size of its relative, the large
coelophysoid Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
''
Dilophosaurus ''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 193 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserved ...
''. The size of its
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
was 469 mm, which was comparable to that of ''
Liliensternus ''Liliensternus'' is an extinct genus of basal neotheropod dinosaur that lived approximately 210 million years ago during the latter part of the Triassic Period in what is now Germany. ''Liliensternus'' was a moderate-sized, bipedal, ground-dwel ...
'' (409 mm) and ''
Dilophosaurus ''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 193 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserved ...
'' (555 mm), suggesting that it was a large Triassic dinosaur, estimated to be about 5.5 m (18 ft) long, which can be extrapolated to a weight of approximately 150–200 kg (330–440 lb). Carpenter (1997) pointed to features of the pelvis and ankle suggesting that this was an immature individual, and could therefore have grown to even a larger size in maturity. Christopher T. Griffith (2019) confirmed ''Gojirasaurus'' possessed features indicative of ontogenetic immaturity. Specimen NMMNH P-4666, which consists of only a pubis, was referred to this genus by Hunt in 1994.


Classification

In 1994 Adrian Hunt, in his unpublished thesis, described and named this material "Revueltoraptor lucasi" which is now considered a
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
. Carpenter officially described and named UCM 47221, ''Gojirasaurus quayi'' in 1997 and classified it as a
coelophysoid Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
. The original remains attributed to this dinosaur included a serrated tooth, a cervical rib, two anterior dorsal ribs, one posterior dorsal rib, a right scapula, two gastralia, four
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, one anterior chevron, a right pubis, a left
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
, and one metatarsal. Tykoski and Rowe (2004) and later Carrano ''et al.'' (2005) agreed that ''Gojirasaurus'' is more derived than ''
Dilophosaurus ''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 193 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserved ...
'' However, later study by Nesbitt ''et al.'' (2007) showed that the vertebrae actually belonged to the
rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia ...
n ''
Shuvosaurus ''Shuvosaurus'' (meaning "Shuvo's lizard") is a genus of beaked reptile from the Late Triassic of western Texas. Despite looking superficially similar to a theropod dinosaur, it is actually more closely related to crocodilians. Discovery and cl ...
'', and the pubis and
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
belonged to another
coelophysoid Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
, indistinguishable from the contemporary ''Coelophysis'', making the status of ''Gojirasaurus'' as a valid genus dubious.


Distinguishing anatomical features

According to Nesbitt ''et al.'' (2007), ''Gojirasaurus'' can be distinguished based on the fact that its
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
is more robust than that of its relative ''
Coelophysis ''Coelophysis'' ( traditionally; or , as heard more commonly in recent decades) is an extinct genus of coelophysid theropod dinosaur that lived approximately 228 to 201.3 million years ago during the latter part of the Triassic Period from t ...
''. Mortimer (2012) has proposed that the observed difference in the leg bone might be just size-related, and not a true
apomorphy In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
. Rauhut (2003) attempted to diagnose this genus based on the fact that the mid/posterior dorsal
vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...
had taller neural spines than those observed in other
coelophysoids Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
. However, the reassignment of the dorsal vertebrae on which the diagnosis was based would render it invalid.


Paleoecology

The only known specimen of ''Gojirasaurus'' was discovered in the
Cooper Canyon Formation The Cooper Canyon Formation is a geological formation of Norian age in Texas and New Mexico.Weishampel, David B; et al. (2004).Dinosaur distribution (Late Triassic, North America)" In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (e ...
of the
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Color ...
near Revuelto Creek, Quay County, in
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
. This genus was discovered in 1981, in gray carbonaceous mudstone deposited during the
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
stage of the Late Triassic, which based on magnetostratigraphy,Molina-Garza, R. S., Geissman, J. W., and Lucas, S. G. 1993. Late Carnian-early Norian magnetostratigraphy from nonmarine strata, Chinle Group, New Mexico. was approximately 228 to 208 million years ago. This specimen is housed in the collection of the University of Colorado Museum, in Boulder, Colorado. ''Gojirasauruss assignment to the
Coelophysoidea Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the ...
, would suggest that it was a bipedal, terrestrial, actively mobile carnivore. Contemporaries of ''Gojirasaurus'' included the pseudosuchian ''
Shuvosaurus ''Shuvosaurus'' (meaning "Shuvo's lizard") is a genus of beaked reptile from the Late Triassic of western Texas. Despite looking superficially similar to a theropod dinosaur, it is actually more closely related to crocodilians. Discovery and cl ...
'', and the phytosaur ''
Rutiodon ''Rutiodon'' ("Wrinkle tooth") is an extinct genus of phytosaur belonging to the family Parasuchidae. It lived during the Late Triassic period, and was about in length. ''Rutiodon'' is known from the eastern United States (North Carolina, New J ...
''.


See also

* Timeline of coelophysoid research


References


External links


Specimen NMMNH P-4666
{{Taxonbar, from=Q134199 Late Triassic dinosaurs of North America Coelophysoids Godzilla (franchise) Taxa named by Kenneth Carpenter Paleontology in New Mexico