Gobiosuchidae
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Gobiosuchidae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
crocodyliforms Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseu ...
known from
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 millio ...
and
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
.


Genera

Three genera are currently classified within Gobiosuchidae: ''
Cassissuchus ''Cassissuchus'' is an Extinction, extinct genus of Gobiosuchidae, gobiosuchid Crocodyliformes, crocodyliform known from the Early Cretaceous Calizas de La Huérgina Formation in Spain. It contains a single species, ''Cassissuchus sanziuami''. ...
'', '' Gobiosuchus'', and '' Zaraasuchus''.Ángela D. Buscalioni (2017) The Gobiosuchidae in the early evolution of Crocodyliformes. Journal of Vertebrate Paleontology Article: e1324459 doi: https://dx.doi.org/10.1080/02724634.2017.1324459 http://www.tandfonline.com/doi/full/10.1080/02724634.2017.1324459


Synapomorphies

According to Pol & Norell (2004), gobiosuchids form a clade united by the following
synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...
: * Parietal without broad occipital portion * Absence of external mandibular fenestra * More than two parallel rows of dorsal
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinc ...
s * Cranial table as wide as ventral portion of skull *
Palpebral An eyelid is a thin fold of skin that covers and protects an eye Eyes are organs of the visual system. They provide living organisms with vision, the ability to receive and process visual detail, as well as enabling several photo respon ...
s sutured to each other and the frontal, excluding it from the orbital margin * External surface of ascending process of
jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anat ...
exposed posterolaterally * Longitudinal ridge on lateral surface of jugal below infratemporal fenestra * Dorsal surface of posterolateral process of
squamosal The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal bones form the cheek series of the skull. The bone forms an ancestral co ...
ornamented with three longitudinal ridges * Presence of a sharp ridge along ventral surface of angular *
Surangular The suprangular or surangular is a jaw bone found in most land vertebrates, except mammals. Usually in the back of the jaw, on the upper edge, it is connected to all other jaw bones: dentary, angular, splenial and articular The articular bone ...
with a longitudinal ridge on its dorsolateral surface * Dorsal surface of osteoderms ornamented with anterolaterally and anteromedially directed ridges * Cervical region surrounded by lateral and ventral osteoderms sutured to the dorsal elements * Presence of appendicular osteoderms * Closed, or incipiently close, supratemporal fenestra


Sources

* Pol, D. & Norell, M. A., (2004). "A new gobiosuchid crocodyliform taxon from the Cretaceous of Mongolia". ''American Museum Novitates'' 3458: 1-31.


References

{{Taxonbar, from=Q1012863 Early Cretaceous crocodylomorphs Late Cretaceous crocodylomorphs Terrestrial crocodylomorphs Early Cretaceous first appearances Late Cretaceous extinctions Prehistoric reptile families