Glochidion Wrightii
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''Glochidion'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of flowering plants, of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Phyllanthaceae Phyllanthaceae is a family of flowering plants in the eudicot order Malpighiales. It is most closely related to the family Picrodendraceae.Kenneth J. Wurdack and Charles C. Davis. 2009. "Malpighiales phylogenetics: Gaining ground on one of the ...
, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, distributed from
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
to the
Pacific Islands Collectively called the Pacific Islands, the islands in the Pacific Ocean are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of se ...
. ''Glochidion'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including ''
Aenetus eximia ''Aenetus eximia'' is a moth of the family Hepialidae. It is known from southern Queensland, Australia, to Tasmania. The wingspan is about 70 mm. Adult males are blue-green. Females are green with two spots on each forewing. The hindwings ar ...
'' and ''
Endoclita damor ''Endoclita damor'' is a species of moth of the family Hepialidae. It is known from India and the Himalayas. Food plants for this species include ''Albizia'', ''Altingia'', ''Cinchona'', ''Coffea'', ''Erythrina'', ''Eugenia'', ''Glochidion'', ''Ma ...
''. The
Nicobarese people The Nicobarese people are an Austroasiatic-speaking people of the Nicobar Islands, a chain of islands in the Bay of Bengal north of Sumatra, forming part of the union territory of Andaman and Nicobar Islands, India. Only 12 of the 19 islands are ...
have attested to the medicinal properties found in ''G. calocarpum'', saying that its bark and seed are most effective in curing abdominal disorders associated with
amoebiasis Amoebiasis, or amoebic dysentery, is an infection of the intestines caused by a parasitic amoeba ''Entamoeba histolytica''. Amoebiasis can be present with no, mild, or severe symptoms. Symptoms may include lethargy, loss of weight, colonic ulce ...
. ''Glochidion'' are of note in the fields of
pollination biology Anthecology, or pollination biology, is the study of pollination as well as the relationships between flowers and their pollinators. Floral biology is a bigger field that includes these studies. Most flowering plants, or angiosperms, are pollinate ...
and
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
because they have a specialized mutualism with moths in the genus ''
Epicephala ''Epicephala'' (leafflower moths) is a genus of moths in the family Gracillariidae. ''Epicephala'' is of note in the fields of pollination biology and coevolution because many species in this genus are pollinators of plants in the genera ''Glo ...
'' (leafflower moths), in which the moths actively pollinate the flowers—thereby ensuring that the tree may produce viable seeds—but also lay eggs in the flowers' ovaries, where their larvae consume a subset of the developing seeds as nourishment. Other species of ''Epicephala'' are pollinators, and in some cases, non-pollinating seed predators, of certain species of plants in the genera ''
Phyllanthus ''Phyllanthus'' is the largest genus in the plant family Phyllanthaceae. Estimates of the number of species in this genus vary widely, from 750David J. Mabberley. 2008. ''Mabberley's Plant-Book.'' third edition (2008). Cambridge University P ...
'' and ''
Breynia ''Breynia'' is a plant genus in the family Phyllanthaceae, first described in 1776. It is native to Southeast Asia, China, the Indian Subcontinent, Papuasia, Australia, and the island of Réunion. The name ''Breynia'' is a conserved name, it ...
'', both closely related to ''Glochidion''. This relationship is similar to those between
figs The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
and
fig wasp Fig wasps are wasps of the superfamily Chalcidoidea which spend their larval stage inside figs. Most are pollinators but others simply feed off the plant. The non-pollinators belong to several groups within the superfamily Chalcidoidea, while th ...
s and
yucca ''Yucca'' is a genus of perennial shrubs and trees in the family Asparagaceae, subfamily Agavoideae. Its 40–50 species are notable for their rosettes of evergreen, tough, sword-shaped leaves and large terminal panicles of white or whitish flo ...
s and yucca moths. Although the genus ''Glochidion'' is native only to the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
, the East Asian species ''
Glochidion puberum ''Glochidion puberum'' is a species of shrub or small tree in the family Phyllanthaceae. It is native to China, where it is widely distributed in both subtropical and temperate regions (Anhui, Fujian, Gansu, Guangdong, Guizhou, Hainan, Henan, ...
'' has become naturalized at several locations in the U.S. state of
Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...
. In a 2006 revision of the Phyllanthaceae, it was recommended that ''Glochidion'' be subsumed in ''Phyllanthus''. New combinations in ''Phyllanthus'' have been published for Madagascar and the Pacific Islands,Wagner WL, Lorence DH. A nomenclator of Pacific oceanic island Phyllanthus (Phyllanthaceae), including Glochidion. PhytoKeys. 2011(4):67-94 but most remain to be published.


Selected species

An incomplete listing: *''
Glochidion apodogynum ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the ...
'' *''
Glochidion barronense ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the ...
'' *''
Glochidion benthamianum ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the ...
'' *'' Glochidion bourdillonii'' *'' Glochidion calocarpum'' *''
Glochidion christophersenii ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the P ...
'' *''
Glochidion carrickii ''Glochidion carrickii'' is a species of plant in the family Phyllanthaceae. It is a tree endemism, endemic to Peninsular Malaysia. It is threatened by habitat loss. References

Glochidion, carrickii Endemic flora of Peninsular Malaysia ...
'' *'' Glochidion comitum'' *'' Glochidion disparipes'' *''
Glochidion eriocarpum ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the P ...
'' *''
Glochidion ferdinandi ''Glochidion ferdinandi'', with common names that include cheese tree (see below), is a species of small to medium–sized trees, constituting part of the plant family Phyllanthaceae. They grow naturally across eastern Australia, from south–eas ...
'' - cheese tree, buttonwood, pencil cedar, water gum, rain tree :* var. ''ferdinandi'' :* var. ''pubens'' *''
Glochidion gardneri ''Glochidion gardneri'' is a species of plant in the family Phyllanthaceae. It is endemic to Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-s ...
'' *'' Glochidion grantii'' *'' Glochidion harveyanum'' - buttonwood, daphne buttonwood :* var. ''harveyanum'' :* var. ''pubescens'' *''
Glochidion hylandii ''Glochidion hylandii '' known as the buttonwood, Hyland's buttonwood or pin flower tree is a plant in the family Phyllanthaceae. It is found in tropical north eastern Australia in the vicinity of the Atherton Tableland. Usually a small tree foun ...
'' *'' Glochidion insulare'' *'' Glochidion johnstonei'' *''
Glochidion lanceolarium ''Glochidion lanceolarium'' is a species of leafflower tree in the family Phyllanthaceae. It is native to Cambodia, China (Fujian, Guangdong, Guangxi, Hainan, and Yunnan), India, Laos, Thailand, and Vietnam. In Mandarin it is known as 艾胶 ...
'' *'' Glochidion lobocarpum'' *''
Glochidion longfieldiae ''Glochidion longfieldiae'', or kaema in the Rapa language, is a species of tree or shrub in the family Phyllanthaceae. It is endemic to the island of Rapa in the Austral Islands in French Polynesia. Molecular phylogenetic analysis indicates tha ...
'' *''
Glochidion manono ''Glochidion manono'', also known as manono or mahame in Tahitian, is a species of shrub or small tree in the family Phyllanthaceae. It is endemic to the Windward Society Islands in French Polynesia, where it is found on the islands of Tahiti ...
'' *'' Glochidion marchionicum'' *'' Glochidion moonii'' *'' Glochidion moorei'' *''
Glochidion myrtifolium ''Phyllanthus st-johnii'' is a species of tree or shrub in the family Phyllanthaceae. It is endemic to the Society Islands of French Polynesia, where it is native to the islands of Bora Bora, Tahaa, Raiatea, and Moorea Moorea ( or ; Tahitia ...
'' *''
Glochidion nadeaudii ''Glochidion nadeaudii'' is a species of tree in the family Phyllanthaceae. It is endemic to the island of Mo'orea in French Polynesia )Territorial motto: ( en, "Great Tahiti of the Golden Haze") , anthem = , song_type = Regional anthem , s ...
'' *''
Glochidion orientalis ''Glochidion'' is a genus of flowering plants, of the family (biology), family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from M ...
'' *'' Glochidion papenooense'' *''
Glochidion pauciflorum ''Glochidion pauciflorum'' is a species of plant in the family Phyllanthaceae. It is native to Karnataka and Tamil Nadu Tamil Nadu (; , TN) is a state in southern India. It is the tenth largest Indian state by area and the sixth larges ...
'' *'' Glochidion perakense'' - umbrella cheese tree, buttonwood :* var. ''supra-axillare'' *'' Glochidion philippicum'' *'' Glochidion pitcairnense'' *'' Glochidion pruinosum'' *''
Glochidion puberum ''Glochidion puberum'' is a species of shrub or small tree in the family Phyllanthaceae. It is native to China, where it is widely distributed in both subtropical and temperate regions (Anhui, Fujian, Gansu, Guangdong, Guizhou, Hainan, Henan, ...
'' *''
Glochidion pungens ''Glochidion'' is a genus of flowering plants, of the family (biology), family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from M ...
'' *'' Glochidion raivavense'' *''
Glochidion ramiflorum ''Glochidion'' is a genus of flowering plants, of the family (biology), family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from M ...
'' J.R.Forst. & G.Forst. *''
Glochidion rapaense ''Glochidion rapaense'', also known by its synonym ''Phyllanthus rapaense'', is a species of tree in the family Phyllanthaceae. It is endemic to the island of Rapa in the Austral Islands of French Polynesia )Territorial motto: ( en, "Great Tah ...
'' *''
Glochidion seemannii ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the P ...
'' Müll.-Arg. *'' Glochidion sessiliflorum'' :* var. ''pedicellatum'' :* var. ''sessiliflorum'' :* var. ''stylosum'' *'' Glochidion sisparense'' *''
Glochidion stellatum ''Glochidion stellatum'' is a species of plant in the family Phyllanthaceae. It is endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zo ...
'' *'' Glochidion stylosum'' *''
Glochidion sumatranum ''Glochidion sumatranum '' known as the umbrella cheese tree is a plant in the family Phyllanthaceae. Although recognized as a distinct species by some sources, others include it within ''Glochidion zeylanicum''. It is found in northern and easte ...
'' *''
Glochidion symingtonii ''Glochidion symingtonii'' is a species of plant in the family Phyllanthaceae. It is a tree endemic to Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Nege ...
'' *''
Glochidion taitense ''Glochidion taitense'', also known by the synonym ''Phyllanthus taitensis'' or as mahame in Tahitian, is a species of tree or shrub in the family Phyllanthaceae. It is endemic to the Windward Society Islands of French Polynesia.Florence, J. (1 ...
'' *'' Glochidion temehaniense'' *'' Glochidion tomentosum'' *'' Glochidion tooviianum'' *''
Glochidion xerocarpum ''Glochidion'' is a genus of flowering plants, of the family (biology), family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from M ...
''


References


External links


Youtube video of a ''Glochidion lanceolarium'' flower being pollinated by an ''Epicephala lanceolaria'' moth
{{Taxonbar, from=Q1245856 Phyllanthaceae genera