Glischrochilus Quadripunctatus 2
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''Glischrochilus'' (sometimes misspelled as ''Glisrochilus'') is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of sap-feeding and predatory
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s under the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Nitidulidae The sap beetles, also known as Nitidulidae, are a family (biology), family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antenna (biology), antennae. Some have red or yellow spots or bands. They fe ...
, subfamily
Cryptarchinae Cryptarchinae is a subfamily of sap-feeding beetles in the family Nitidulidae. There are at least 3 genera and about 17 described species in Cryptarchinae. Genera These three genera belong to the subfamily Cryptarchinae: * ''Cryptarcha'' Shucka ...
. Most members of this genus are commonly known as picnic beetles or beer bugs.


Description

''Glischrochilus'' are
oblong An oblong is a non-square rectangle. Oblong may also refer to: Places * Oblong, Illinois, a village in the United States * Oblong Township, Crawford County, Illinois, United States * A strip of land on the New York-Connecticut border in the Unit ...
shiny black beetles with attractive yellow, red, or orange markings on their
elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...
. Their elytra are short and expose the upper surface of their last abdominal segments, a good way to distinguish them from the superficially similar but generally larger '' Megalodacne'' beetles. They are so similar that some
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''Glischrochilus'' were once classified along with ''Megalodacne'' under the now reclassified genus ''
Ips IPS, ips, or iPS may refer to: Science and technology Biology and medicine * ''Ips'' (genus), a genus of bark beetle * Induced pluripotent stem cell or iPS cells * Intermittent photic stimulation, a neuroimaging technique * Intraparietal sulcus, ...
''. Like other nitulidid beetles, adult ''Glischrochilus'' can be distinguished from other kinds of sap-feeding beetles by their characteristic 11-segmented antennae that end with a 3-segmented ball-like club. ''Glischrochilus'' are among the largest of the nitulidid beetles, but they are still generally smaller in comparison to other beetles, averaging at only in length, with larger specimens at long. ''Glischrochilus'' eggs are sausage-shaped and milky white. Eggs are laid during
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a ...
near decaying plant matter.
Larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e are about long and feed for three weeks on fermenting juices and then
pupate A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
. It takes a little over a month for picnic beetles to develop from egg to adult and only one generation is produced each year.


Ecology

''Glischrochilus'' beetles from the subgenus ''Librodor'', consisting the majority of species in the genus, feed on exuding sap from injured trees and decaying vegetable or fungal matter. They are also attracted to ripening fruits, as well as
beer Beer is one of the oldest and the most widely consumed type of alcoholic drink in the world, and the third most popular drink overall after water and tea. It is produced by the brewing and fermentation of starches, mainly derived from ce ...
,
vinegar Vinegar is an aqueous solution of acetic acid and trace compounds that may include flavorings. Vinegar typically contains 5–8% acetic acid by volume. Usually, the acetic acid is produced by a double fermentation, converting simple sugars to et ...
,
wine Wine is an alcoholic drink typically made from fermented grapes. Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different varieties of grapes and strains of yeasts are m ...
,
fruit juice Juice is a drink made from the extraction or pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat or seafood, such as ...
and fermenting beverages. They frequently drown as they feed, rendering these liquids unsuitable for consumption. They congregate in large numbers when such beverages are present, often ruining
picnic A picnic is a meal taken outdoors ( ''al fresco'') as part of an excursion, especially in scenic surroundings, such as a park, lakeside, or other place affording an interesting view, or else in conjunction with a public event such as preceding ...
s and outdoor gatherings like
barbecue Barbecue or barbeque (informally BBQ in the UK, US, and Canada, barbie in Australia and braai in South Africa) is a term used with significant regional and national variations to describe various cooking methods that use live fire and smoke t ...
s, earning them their
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
s of 'picnic beetles', 'picnic bugs', or 'beer bugs'. Researchers who wish to attract the bugs use bait that contains beer, molasses, vinegar, pineapple and other ingredients. ''Glischrochilus'' beetles from the subgenus ''Glischrochilus'' on the other hand are facultative and obligatory
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s of soft
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s (including insect larvae) living under tree barks. Species from both subgenera are found in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
. ''Glischrochilus'' are also known to be involved in the transmission of the
plant pathogen Plant pathology (also phytopathology) is the scientific study of diseases in plants caused by pathogens (infectious organisms) and environmental conditions (physiological factors). Organisms that cause infectious disease include fungi, oomyc ...
ic
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
''
Ceratocystis ''Ceratocystis'' is a genus of fungi in the family Ceratocystidaceae. Several species are important plant pathogens, causing diseases such as oak wilt and pineapple black rot. Species *''Ceratocystis acericola'' *''Ceratocystis acoma'' *''Cerat ...
'' and ''
Fusarium ''Fusarium'' is a large genus of filamentous fungi, part of a group often referred to as hyphomycetes, widely distributed in soil and associated with plants. Most species are harmless saprobes, and are relatively abundant members of the soil mi ...
''. They are also considered pests of certain fruit and vegetable crops like
strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
,
corn Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...
,
tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
es,
apricot An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus ''Prunus''. Usually, an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are also ...
,
muskmelon ''Cucumis melo'', also known as melon, is a species of ''Cucumis'' that has been developed into many cultivated varieties. The fruit is a pepo. The flesh is either sweet or bland, with or without a musky aroma, and the rind can be smooth (such as ...
s,
raspberries The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...
, and
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-fu ...
es. They normally only become a problem when fruits are damaged or are overripe and beginning to ferment. They are difficult to control as they are primarily attracted to the odor of food. Methods of control include scent baits and removing damaged or overripe fruits.


Taxonomy

''Glischrochilus'' belongs to the subfamily
Cryptarchinae Cryptarchinae is a subfamily of sap-feeding beetles in the family Nitidulidae. There are at least 3 genera and about 17 described species in Cryptarchinae. Genera These three genera belong to the subfamily Cryptarchinae: * ''Cryptarcha'' Shucka ...
under the tribe Cryptarchini. It contains two
subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
, ''
Glischrochilus ''Glischrochilus'' (sometimes misspelled as ''Glisrochilus'') is a genus of sap-feeding and predatory beetles under the Family (biology), family Nitidulidae, subfamily Cryptarchinae. Most members of this genus are commonly known as picnic beetles ...
'' and '' Librodor''. They were first described by the
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
entomologist Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arach ...
Edmund Reitter Edmund Reitter (22 October 1845 – 15 March 1920) was an Austrian entomologist, writer and a collector. Biography Edmund Reitter was best known as an expert on the beetles of the Palaearctic. He was an imperial advisor and editor of the ...
in 1873. The name ''Glischrochilus'' is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
words ' (sticky) and (', lip).


List of species

The following list may be incomplete or inaccurate: *Subgenus ''Glischrochilus'' Reitter, 1873 ** '' Glischrochilus biguttulus'' (Motschulsky, 1860) ** '' Glischrochilus confluentus'' (Say) ** '' Glischrochilus cruciatus'' (Motschulsky, 1860) ** '' Glischrochilus lecontei'' (Brown, 1931) ** '' Glischrochilus moratus'' (Brown) ** '' Glischrochilus obtusus'' (Say) ** '' Glischrochilus quadripunctatus'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of Systema Naturae, 10th edition of ''Systema Naturae'', the sta ...
)
– European bark beetle predator ** '' Glsichrochilus vittatus'' (Say) *Subgenus ''Librodor'' Reitter, 1884 ** '' Glischrochilus affinis'' Kirejtshuk, 1984 ** '' Glischrochilus binaevus'' (Reitter, 1879) ** '' Glischrochilus christophi'' (Reitter, 1879) ** '' Glischrochilus clavatus'' Reitter, 1884 ** '' Glischrochilus fasciatus'' (Olivier, 1790) – red-spotted sap beetle ** '' Glischrochilus grandis'' (Tournier, 1872) ** '' Glischrochilus hortensis'' (Fourcroy, 1785) ** '' Glischrochilus ipsoides'' (Reitter, 1879) ** '' Glischrochilus japonicus'' (Motschulsky, 1858) ** '' Glischrochilus jelineki'' Lasoń, 2009 ** '' Glischrochilus pantherinus'' (Reitter, 1879) ** '' Glischrochilus parvipustulatus'' (Kolbe, 1886) ** '' Glischrochilus quadriguttatus'' (Fabricius, 1776) ** ''
Glischrochilus quadrisignatus ''Glischrochilus quadrisignatus'', known generally as four-spotted sap beetle, is a species of sap-feeding beetle in the family Nitidulidae. Other common names include the beer bug and picnic beetle. It is found in North America. Four-spotted s ...
'' (Say, 1835) – four-spotted sap beetle or picnic beetle ** '' Glischrochilus rufiventris'' (Reitter, 1879) ** '' Glischrochilus sanguinolentus'' (Olivier, 1790) ** '' Glischrochilus siepmanni'' (W. J. Brown) ** '' Glischrochilus subcylindricus'' Reitter, 1884 * Species '' Glischrochilus becvari'' Jelinek, 1999 * Species '' Glischrochilus forcipatus'' Fairmaire, 1889 * Species '' Glischrochilus vittatus'' (Say, 1835)


References


Notes

: For example
Finsberg, T. A. C. 2009. Insektsliv och vegetation vid brandfältet på Stora Fjället ett år efter, pp.13


External links

* * {{Taxonbar, from=Q3108901 Nitidulidae