''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy areas worldwide (except Australia), only the nocturnal adults have ever been collected (typically at lights), and both the larvae and biology of ''Glaresis'' are as yet unknown. Due to their narrow habitat associations, a great number of these species occur in extremely limited geographic areas, and are accordingly imperiled by habitat destruction. These beetles are small, 2.5–6 mm long, and have the stocky appearance typical of

fossorial A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric eviden ...

scarabs, with short, heavy, spurred legs. Color ranges from tan to dark brown, and the back is covered with short setae. Efforts to raise glaresids in the laboratory were undertaken in the 1980s by

C. H. Scholtz C. or c. may refer to: * Century, sometimes abbreviated as ''c.'' or ''C.'', a period of 100 years * Cent (currency), abbreviated ''c.'' or ''¢'', a monetary unit that equals of the basic unit of many currencies * Caius or Gaius, abbreviated as ...

and others, but were unsuccessful. ''Glaresis'' was originally classified with Trogidae (originally a subfamily within Scarabaeidae), and has many characteristics of "primitive" scarabaeoids, but no affinities to any of the other primitive groups; recent work suggests that they may in fact belong in Trogidae. Scholtz argued that ''Glaresis'' is the most primitive type of scarabaeoid, but more recent research indicates that the Pleocomidae hold this position. The species in North, Central, and South America have been revised by Robert Gordon and Guy Hanley, January 2014, in the journal ''Insecta Mundi''.

Species

* ''

Glaresis alfierii ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis arabica ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis arenata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis australis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * '' Glaresis bajaensis'' * '' Glaresis bautista'' * ''

Glaresis beckeri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis caenulenta ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * '' Glaresis california'' * '' Glaresis canadensis'' * '' Glaresis carthagensis'' * '' Glaresis cartwrighti'' * '' Glaresis ceballosi'' * '' Glaresis celiae'' * '' Glaresis clypeata'' * '' Glaresis confusa'' * '' Glaresis contrerasi'' * '' Glaresis costaricensis'' * ''

Glaresis costata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis dakotensis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis dentata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis desperata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis donaldi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis ecostata ''Glaresis ecostata'' is a species of enigmatic scarab beetle in the family Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was former ...

'' * '' Glaresis exasperata'' * ''

Glaresis falli ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis foveolata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis franzi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis freyi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis fritzi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis frustrata ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis gineri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis gordoni ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis handlirschi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis hespericula ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis hispana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis holmi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis holzschuhi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis howdeni ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis imitator ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis impressicollis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis inducta ''Glaresis inducta'' is a species of enigmatic scarab beetle in the family Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerl ...

'' * ''

Glaresis kocheri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis koenigsbaueri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * '' Glaresis limbata'' * ''

Glaresis lomii ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis longisternum ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * '' Glaresis mandibularis'' * ''

Glaresis maroccana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis mauritanica ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis medialis ''Glaresis medialis'' is a species of enigmatic scarab beetle in the family Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was former ...

'' * ''

Glaresis mendica ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis methneri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis minuta ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis mondacai ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis montenegro ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis namibensis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis obscura ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis ordosensis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis orientalis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis oxiana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis paramendica ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis pardoalcaidei ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis pardoi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis penrithae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis phoenicis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis porrecta ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis quedenfeldti ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis rufa ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis smithi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis subulosa ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis texana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis thiniensis ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis tripolitana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis tumida ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis villiersi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis walzlae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis warneri ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis yanegai ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

'' * ''

Glaresis zacateca ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy a ...

'' * ''

Glaresis zarudniana ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy area ...

'' * ''

Glaresis zvirgzdinsi ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy are ...

Fossil species

* †''Glaresis burmitica'' Cai and Huang 2018 Burmese amber, Myanmar,

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

* †''Glaresis orthochilus'' Bai et al. 2010 Yixian Formation, China,

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

* †''Glaresis tridentata'' Bai et al. 2014M. Bai, R. G. Beutel, and D. Ren. 2014
Description of a new species of Glaresidae (Coleoptera: Scarabaeoidea) from the Jehol Biota of China with a geometric morphometric evaluation.
''Arthropod Systematics & Phylogeny'' 72:223-236 Yixian Formation, China, Aptian ''"Glaresis" cretacea'' Nikolajev 2007 from the Aptian aged Zaza Formation in Russia is considered questionable, with Cai and Huang 2018 considering it to have no diagnostic characters of the family.

References

* Mary Liz Jameson, "Glaresidae", in Ross H. Arnett, Jr. and

Michael C. Thomas Michael Charles Thomas (May 5, 1948 – October 14, 2019) was an American entomologist who co-authored the book series ''American Beetles''. Born in Miami, Florida, Thomas graduated from the University of South Florida in 1970 with a Bachelor o ...

, ''American Beetles'' (

CRC Press The CRC Press, LLC is an American publishing group that specializes in producing technical books. Many of their books relate to engineering, science and mathematics. Their scope also includes books on business, forensics and information tec ...

, 2001), vol. 2

External links

* Browne et al.
''Glaresis''
from th
Tree of Life
{{Taxonbar, from1=Q140720, from2=Q14868999 Scarabaeoidea genera Scarabaeiformia