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A paca is a member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Cuniculus'' of ground-dwelling, herbivorous
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s in South and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
. It is the only genus in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Cuniculidae. Pacas are large rodents with dots and stripes on their sides, short ears, and barely visible tails. Pacas are eaten by people in Belize, where they are known as "gibnut" and, having been served to Queen Elizabeth II, "the royal rat". In the Amazon basin they are known as "majás". The
Smithsonian Tropical Research Institute The Smithsonian Tropical Research Institute (STRI, es, Instituto Smithsonian de Investigaciones Tropicales) is located in Panama and is the only bureau of the Smithsonian Institution based outside of the United States. It is dedicated to understa ...
in Panama has studied the possibilities of developing the paca as a viable high-priced food supply for the tropics.


Evolutionary background

Pacas originated in South America and are one of the few mammal species that successfully emigrated to North America after the Great American Interchange . They were formerly grouped with the
agoutis The agouti (, ) or common agouti is any of several rodent species of the genus ''Dasyprocta''. They are native to Middle America, northern and central South America, and the southern Lesser Antilles. Some species have also been introduced else ...
in the family
Dasyproctidae Dasyproctidae is a family of large South American rodents, comprising the agoutis and acouchis. Their fur is a reddish or dark colour above, with a paler underside. They are herbivorous, often feeding on ripe fruit that falls from trees. They liv ...
, subfamily Agoutinae, but were given full family status because they differ in the number of toes, the shape of the skull, and coat patterning.


Description

left, Paca Pacas are in length, excluding the short tail, weigh , and are the sixth-largest rodents in the world. Similar to
guinea pig The guinea pig or domestic guinea pig (''Cavia porcellus''), also known as the cavy or domestic cavy (), is a species of rodent belonging to the genus '' Cavia'' in the family Caviidae. Breeders tend to use the word ''cavy'' to describe the ...
s, they have square heads, small ears, sides patterned with spots and stripes, and virtually invisible tails. With large hind limbs, small fore limbs, and cone-shaped bodies, pacas are similar in appearance to the deer-like, ungulate
chevrotain Chevrotains, or mouse-deer, are small even-toed ungulates that make up the family Tragulidae, the only extant members of the infraorder Tragulina. The 10 extant species are placed in three genera, but several species also are known only ...
s, and like them have four to seven horizontal lines of blotches and stripes along their flanks. They have a heavy and robust appearance, though their legs are long and relatively tiny. Their small ears are set high on their heads. They have four toes on their fore feet and five on their hind feet (of which two are short and hardly touch the ground) and they have stout nails that resemble small hooves. In young pacas, the skin is covered with horny scales about in diameter; perhaps these scales have a protective function against smaller predators. There is virtually no difference between sexes. They can live up to 13 years in the wild.


Behavior

Pacas inhabit rainforests, cloud forests, and sometimes more open habitats. They are great swimmers and prefer to be near water. They dive when threatened and can stay submerged up to 15 minutes. They can also jump up to and freeze for up to 45 minutes. They normally move along well-established paths and will create new paths when old ones are disturbed. They are normally passive in daytime and forage in the morning and afternoon, but can be strictly nocturnal in areas with many predators. They live in burrows up to deep, normally with two entrances covered with leaves to hide the burrow and to serve as an early warning system. Burrows are often near water, but always above the seasonal flood line. Predators except humans include jaguar, puma, ocelot,
margay The margay (''Leopardus wiedii'') is a small wild cat native to Central and South America. A solitary and nocturnal cat, it lives mainly in primary evergreen and deciduous forest. Until the 1990s, margays were hunted illegally for the wildlif ...
, jaguarundi,
bush dog The bush dog (''Speothos venaticus'') is a canine found in Central and South America. In spite of its extensive range, it is very rare in most areas except in Suriname, Guyana and Peru; it was first identified by Peter Wilhelm Lund from fossi ...
,
boa constrictor The boa constrictor (scientific name also ''Boa constrictor''), also called the red-tailed boa, is a species of large, non-venomous, heavy-bodied snake that is frequently kept and bred in captivity. The boa constrictor is a member of the family B ...
, and
caiman A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America f ...
. Pacas have resonating chambers in their cheeks and their growling noise, at about 1 kHz, is surprisingly loud for their size. Aside from making noises, territories are marked with urine. Population density can reach up to 70 adults per , and pacas often constitute some 20% of the biomass of terrestrial mammals.


Feeding

In the wild, pacas eat fruits from
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
trees and fallen fruits from taller trees, but may also eat leaves, buds, flowers, fungi, and insects. They play a vital role in seed dispersal, and, with seasonal adaptations, their home ranges are often centered on a group of fruit trees. Pacas normally do not use their fore-paws to manipulate fruits (as do agoutis), instead using their powerful jaw muscles to break open hard-shelled fruits. Unlike agoutis, pacas can store fat, and so are less dependent on the caching of seeds. Competition from agoutis is avoided by a slight variation in activity cycles and food preferences. Like rabbits, pacas are
coprophagous Coprophagia () or coprophagy () is the consumption of feces. The word is derived from the grc, κόπρος , "feces" and , "to eat". Coprophagy refers to many kinds of feces-eating, including eating feces of other species (heterospecifics), of ...
and absorb protein and carbohydrates from specially produced moist fecal pellets. Before allowing their young to suckle, mothers lick them to stimulate them to defecate and urinate, and then lick the resulting product, both to feed herself and to prevent the odour from attracting predators.


Reproduction

Gestation lasts between 114 and 119 days with about 190 days between births. Pacas are
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
, the young are born with fur and open eyes. Normally, mothers give birth to one young, but she can give birth up to three times per year if conditions allow. More than one birth per year results in lactation periods overlapping pregnancies.
Weaning Weaning is the process of gradually introducing an infant human or another mammal to what will be its adult diet while withdrawing the supply of its mother's milk. The process takes place only in mammals, as only mammals produce milk. The infan ...
begins after six weeks, but the young start to follow their mothers early and can do so for up to a year. Sexual maturity is reached after 6–12 months, when females weigh about and males . Pacas usually mate in water. As the male approaches the female, she starts to hop enthusiastically, more so if he sprays her with urine. Pacas have altered the common rodent strategy — safety in numbers — and mind their offspring carefully. At at birth, the young are born in holes too small for both predators and the mother to enter, which are then covered with leaves and twigs. To invite the young out of the hole, the mother uses a low rolling vocalization. Suckling usually lasts for 90 days, after which the young weighs .


Distribution and habitat

The lowland paca is found from southern Mexico to northern Argentina. They primarily live in rainforests near streams, but can also be found in a wide variety of habitats, including mangrove swamps,
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
s near water currents, and even in public parks. They have been observed up to above sea level. The smaller mountain paca lives in the northern
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
and the Páramo grasslands, with a peak occurrence between above sea level.


Species

The mountain paca has longer and darker fur than the lowland paca. Observations indicate mountain pacas are found between above sea level. *
Lowland paca The lowland paca (''Cuniculus paca''), also known as the spotted paca, is a large rodent found in tropical and sub-tropical America, from east-central Mexico to northern Argentina, and has been introduced to Cuba and Algeria. The animal is cal ...
, ''Cuniculus paca'' ( Agouti paca) * Mountain paca, ''Cuniculus taczanowskii'' * '' Cuniculus hernandezi'' (a recent and disputed '' nomen nudum'')


Citations


General sources

* * * * * *


External links


Gibnuts
at Crooked Tree Wildlife Sanctuary {{Taxonbar, from=Q202697 Hystricognath rodents Mammals of Guyana Coprophagous animals Rodents of Central America Taxa named by Mathurin Jacques Brisson