The giant pika or Wharton's pika (''Ochotona whartoni'') is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

mammal species in the family Ochotonidae. It lived during the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

and early

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...

in northern parts of

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

(

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

, US and

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

). Very similar forms have also been found also in

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

Distribution

The giant pika has been found in Alaska (United States),

Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...

(''O. whartoni'' and ''O''

cf. The abbreviation ''cf.'' (short for the la, confer/conferatur, both meaning "compare") is used in writing to refer the reader to other material to make a comparison with the topic being discussed. Style guides recommend that ''cf.'' be used onl ...

''whartoni'', large number of locations),

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

and

Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...

(Canada). A close relative ''O. whartoni'' (''O.'' cf. ''whartoni'') is also known from Eastern

and

Kolyma Kolyma (russian: Колыма́, ) is a region located in the Russian Far East. It is bounded to the north by the East Siberian Sea and the Arctic Ocean, and by the Sea of Okhotsk to the south. The region gets its name from the Kolyma River an ...

. The ancestors of these pikas migrated from

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

during the Early Pleistocene via the

Bering Land Bridge Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of ...

, along with another group of small pikas close to the "'' O. pusilla'' group". This migration was separate from that of '' O. spanglei'', which entered North America approximately three million earlier at the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58

Detailed fossil distribution

Old Crow River Old Crow River is a transnational stream, long, that begins in the U.S. state of Alaska and flows generally southeast to meet the Porcupine River in the Canadian territory of Yukon. In turn, the Porcupine, a tributary of the Yukon River, flows ba ...

, Irvingtonian (1.8 - 0.3 Ma) and Middle

(0.8 - 0.1 Ma), ''O''

''whartoni'' and ''O. whartoni''. **Old Crow River, Yukon, Late/Upper

(Late

Illinoian The Illinoian Stage is the name used by Quaternary geologists in North America to designate the period c.191,000 to c.130,000 years ago, during the middle Pleistocene, when sediments comprising the Illinoian Glacial Lobe were deposited. It precedes ...

- Sangamonian stages)
(0.1 - 0.0 Ma / ~125,000-80,000 BP), at least 10 locations, ''O. cf. whartoni'' **Thistle Creek, Yukon, Middle Pleistocene (0.8 - 0.1 Ma), ''O. whartoni'' **Eagle Cave,

, >33,000 BP **Elba Cave at

Elba Elba ( it, isola d'Elba, ; la, Ilva) is a Mediterranean island in Tuscany, Italy, from the coastal town of Piombino on the Italian mainland, and the largest island of the Tuscan Archipelago. It is also part of the Arcipelago Toscano National ...

, 8670±220 BP (the last known occurrence of this species) **Kelso Cave,

, Late

Wisconsian The Wisconsin Glacial Episode, also called the Wisconsin glaciation, was the most recent glacial period of the North American ice sheet complex. This advance included the Cordilleran Ice Sheet, which nucleated in the northern North American Cor ...

(>32,000 BP) ::The large form of ''Ochotona'' was found in 2 of 5 localities in eastern

. *

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

** Cape Deceit,

, Cape Deceit Formation, Irvingtonian (1.8 - 0.3 Ma), ''O. whartoni'',
(the species was discovered here) **Gold Hill Cut at

Fairbanks Fairbanks is a home rule city and the borough seat of the Fairbanks North Star Borough in the U.S. state of Alaska. Fairbanks is the largest city in the Interior region of Alaska and the second largest in the state. The 2020 Census put the po ...

, Alaska *

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

- only ''O''

''whartoni'' **From eastern

- Zayarsk site to the

Range area (Krestovka Sections,

Yakutia Sakha, officially the Republic of Sakha (Yakutia),, is the largest republic of Russia, located in the Russian Far East, along the Arctic Ocean, with a population of roughly 1 million. Sakha comprises half of the area of its governing Far Eas ...

), in Kolyma from Late

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...

**Western Siberia, but mentions only ''Ochotona'' as occurring there.

Biology

The giant pika is much larger than other

pika A pika ( or ; archaically spelled pica) is a small, mountain-dwelling mammal found in Asia and North America. With short limbs, very round body, an even coat of fur, and no external tail, they resemble their close relative, the rabbit, but wi ...

s, but is of a similar size to the extinct early and middle Pleistocene ''O. complicidens'' and

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

''O. koslowi'' ( Koslov's pika), both from

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

, and may belong to one of them. Unlike the

American pika The American pika (''Ochotona princeps''), a diurnality, diurnal species of pika, is found in the mountains of western North America, usually in boulder fields at or above the tree line. They are herbivorous, smaller relatives of rabbits and ha ...

(''O. princeps''), which inhabits scree slopes, the giant pika's habitat was largely

tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...

and

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...

, similar to Eurasian pikas.

Occurrence and extinction

The giant pika has been found in North America from the Irvingtonian (1.8–0.3 Ma,

Lower Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...

–

Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. The ...

) throughout Middle Pleistocene to

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...

(0.1–0.0Ma) locations. The last occurrence of the giant pika is known from early the

of eastern North America (a

cave A cave or cavern is a natural void in the ground, specifically a space large enough for a human to enter. Caves often form by the weathering of rock and often extend deep underground. The word ''cave'' can refer to smaller openings such as sea ...

in the

Niagara Escarpment The Niagara Escarpment is a long escarpment, or cuesta, in Canada and the United States that runs predominantly east–west from New York through Ontario, Michigan, Wisconsin, and into Illinois. The escarpment is most famous as the cliff over ...

) and its radiometric date is 8670±220 years BP ( ¹⁴C age) or 10251-9140 BP (calibrated date). It is possible that it survived in the rocky areas along the Niagara Escarpment as a

relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...

population.

Notes

References

Additional references of the Paleobiology Database

{{Taxonbar, from=Q16441259 Pikas Holocene extinctions Prehistoric lagomorphs Extinct animals of the United States Extinct animals of Canada Prehistoric mammals of North America Fossil taxa described in 1971