Geosesarma Dennerle
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''Geosesarma dennerle'' is a species of small land-living
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
s found on
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. It is popular in the aquarium trade, where G. dennerle, in particular, is often simply called "Vampire Crab". Crabs called "Geosesarma bicolor Krakatau Vampirkrabbe" are probably also ''G. dennerle''. All species of '' Geosesarma'' crabs are often called "vampire crabs" in the aquarium trade. ''Geosesarma dennerle'' is, notably, seemingly more carnivorous than other species of ''Geosesarma'', rarely feeding on plant material or decaying plant material The species is named after the aquarium supply company Dennerle, which supported one of the describing authors' (Christian Lukhaup) study in Java. The coloration of ''G. dennerle'' can be very similar to '' G. bicolor''. The describing paper mentions
cheliped A chela ()also called a claw, nipper, or pinceris a pincer-like organ at the end of certain limbs of some arthropods. The name comes from Ancient Greek , through New Latin '. The plural form is chelae. Legs bearing a chela are called chelipeds. ...
s, male abdominal and G1 structures as distinguishing features.


Description

''Geosesarma'' ''dennerle'' is a member of the phylum
Arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
a, and subphylum
Crustacea Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean gro ...
. This species resides in the genus ''Geosesarma de Man'', which comprises 56 recognized species, commonly found in Southeast Asia. These crabs are generally between 1.5-2” in size. The crab features a two-colored
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
, being predominantly purple with a patch of cream/yellow on its back. ''Dennerle'' also possesses bright yellow eyes and vibrant violet claws. As a member of the order
Decapoda The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is esti ...
, this crab has 5 pairs of segmented limbs. Unlike some species of crab that have specialized claws, d''ennerle'' does not have swollen chellae. The carapace is described as square-esque in shape, with well defined regions of the shell, and the abdomen being broad. The ambulatory legs of this crab are generally long and thin, with broad segments. Dennerle often have tubercles along its dorsal carapace, providing some texture to the otherwise smooth shell. ''G. dennerle'' are similar to other species of crabs with the occurrence of
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
. The sex of ''Geosesarma'' species can be identified by observation of abdominal flaps. The males have narrow and pointed abdominal flaps, while the females possess wider flaps which are ovular in shape. Males are also noted to be slightly larger than females.


Ecology

''G.'' ''dennerle'' can be found natively on small islands in the Indian Ocean, such as Java, Sulawesi, Riau, and Krakatau, as well as throughout Asia. As the prefix “geo-” suggests, ''G. dennerle'' is a
terrestrial crab A number of lineages of crabs have evolved to live predominantly on land. Examples of terrestrial crabs are found in the families Gecarcinidae and Gecarcinucidae, as well as in selected genera from other families, such as ''Sesarma'', althoug ...
. It is common to find ''dennerle'' hiding in burrows in areas composed of muddy creek valleys, with rocks and dense vegetation. The diet of ''dennerle'' is greatly composed of insects and plant matter. The high endemism of species within the ''Geosesarma'' genus, means there is not a great amount of geographic distribution.


Reproduction

Sexual reproduction in ''Geosesarma dennerle'' is similar to that of the majority of other species of crabs. Little has been observed towards specific variations in this species reproduction method in comparison to other species in the ''Geosesarma'' genus. Members of the ''G. de Man'' genus commonly produce large eggs, averaging between 1.2 and 1.8 mm in diameter. Members of Geosesarma become sexually mature around 6 months of age, with breeding consisting of the male mounting the female to fertilize eggs. This species has
direct development Marine larval ecology is the study of the factors influencing dispersing larvae, which many marine invertebrates and fishes have. Marine animals with a larva typically release many larvae into the water column, where the larvae develop before metam ...
offspring: fully formed miniature versions of the adults hatch from the eggs. The female will carry around 20-80 fertilized eggs for about a month before hatching occurs. These newly hatched offspring are often independent, and will disperse from the point where they hatch.


Aquatic care

''G. dennerle'', is popular in the aquarium industry for their bright coloration, and is part of a group of crabs nicknamed "Vampire Crabs". The lifespan of Vampire Crabs is said to be around two years in captivity or in the wild. These crabs require freshwater tanks that are at least 10 gallons in volume. Vampire Crabs are sensitive to radical changes in pH and water temperature. They require water that is close to a pH of 7 or 8, with water temperatures in the range of 70-82° Fahrenheit. There are important temperaments of ''Geosesarma'' to understand in order to properly take care of them as pets. They are mostly nocturnal, remaining largely inactive during the day and becoming active at night time. In their tank these crabs prefer having hiding places, as they molt frequently early in their lifespan to grow. Vampire crabs are able to coexist with other species of crabs, but can be aggressive to crabs that are members of the same species or genus. Tank mates such as fish or other invertebrates are recommended to be of similar size, to prevent consumption of either the Vampire crabs or their tank mates.


Breeding in captivity

Currently, there isn't any recommended technique to encourage breeding in captivity, as the behaviors of ''G. dennerle'' are widely unknown. It is recommended that new offspring, when hatched, are to be separated from the adults to prevent cannibalism. If new offspring are kept in the same tank, hiding places are recommended for them to increase survival rate.


References


External links

*
New species of “vampire crabs” (Geosesarma De Man, 1892) from central Java, Indonesia, and the identity of Sesarma (Geosesarma) nodulifera De Man, 1892 (Crustacea, Brachyura, Thoracotremata, Sesarmidae)
* {{Taxonbar, from=Q19345966 Grapsoidea Freshwater crustaceans of Asia Crustaceans described in 2015