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The tokay gecko (''Gekko gecko'') is a
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...
in the genus ''
Gekko ''Gekko'' is a genus of Southeast Asian geckos, commonly known as true geckos or calling geckos, in the family Gekkonidae. Although species such as ''Gekko gecko'' (tokay gecko) are very widespread and common, some species in the same genus hav ...
'', the true geckos. It is native to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
and some
Pacific Islands Collectively called the Pacific Islands, the islands in the Pacific Ocean are further categorized into three major island groups: Melanesia, Micronesia, and Polynesia. Depending on the context, the term ''Pacific Islands'' may refer to one of se ...
.


Subspecies

Two subspecies are currently recognized. *''G. g. gecko'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758): tropical
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
from northeastern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
to eastern
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. *''G. g. azhari'' (
Mertens __NOTOC__ Mertens () is a surname of Flemish Origin, meaning "son of Merten" (Martin). It is the fifth most common name in Belgium with 18,518 people in 2008. Geographical distribution As of 2014, 43.4% of all known bearers of the surname ''Merte ...
, 1955): found only in
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
.


Distribution and habitat

This species is found in northeast India, Bhutan, Nepal, and Bangladesh; throughout Southeast Asia, including Thailand, the Philippines, Malaysia and Indonesia; and toward western New Guinea. Its native habitat is
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
, where it lives on trees and cliffs, and it frequently adapts to rural human habitations, roaming walls and ceilings at night in search of insect prey. This is an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
in some areas outside its native range. It is established in Florida in the United States, Martinique, the islands of Belize, and possibly Hawaii. Increasing urbanization is reducing its range. It is currently unclear whether the species is native but very uncommon in
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
or whether the rare reports of individuals since the 1920s are based on repeated anthropogenic translocations that may or may not have resulted in established populations by now.


Physical characteristics and behavior

The tokay gecko is a large
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
gecko, reaching a total length (including tail) of on average, but some grow as large as long. It is believed to be the third largest species of gecko, after the giant leaf-tail gecko (''Uroplatus giganteus'') and
New Caledonian giant gecko ''Rhacodactylus leachianus'', commonly known as the New Caledonian giant gecko, Leach's giant gecko, Leachianus Gecko, or simply Leachie, is a large species of gecko in the family Diplodactylidae. The species, which was first described by Geor ...
(''Rhacodactylus leachianus''). It is cylindrical but somewhat flattened in body shape. The eyes have vertical pupils. The skin is soft to the touch and is generally blue-gray with red or orange spots and speckles, but the animal can change the color of its skin to blend into the environment. The species is
sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, the males being more brightly colored and slightly larger.Corl, J. 1999
''Gekko gecko''.
Animal Diversity Web. University of Michigan. Retrieved 19 February 2016.
Tokay geckos are generally aggressive, territorial, and can inflict a strong bite. Females lay clutches of one or two hard-shelled eggs and guard them until they hatch. The tokay gecko feeds on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s and small
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
s. In a study conducted in Thailand, researchers noticed that there was little variation in the diets males, females, and juveniles, which was likely due to low insect availability in this area. It is a strong climber with foot pads that can support the entire weight of the body on a vertical surface for a long period of time. Compared to other gecko species, the tokay gecko has a robust build, with a tail, a large head and muscular jaws. Though common in the pet trade, the strong bite of the tokay gecko makes it ill-suited for inexperienced keepers. In addition, the strength of the bite depending on the gecko's size; larger (usually male) tokay geckos are capable of piercing skin, which often results in immediate
bleeding Bleeding, hemorrhage, haemorrhage or blood loss, is blood escaping from the circulatory system from damaged blood vessels. Bleeding can occur internally, or externally either through a natural opening such as the mouth, nose, ear, urethra, vag ...
.


Call

The male's mating call, a loud croak, is variously described as sounding like ''token'', ''gekk-gekk'', ''tuck-too'' or ''tou-kay'' from which both the common and the scientific name (deriving from
onomatopoeic Onomatopoeia is the process of creating a word that phonetically imitates, resembles, or suggests the sound that it describes. Such a word itself is also called an onomatopoeia. Common onomatopoeias include animal noises such as ''oink'', ''m ...
names in
Malay Malay may refer to: Languages * Malay language or Bahasa Melayu, a major Austronesian language spoken in Indonesia, Malaysia, Brunei and Singapore ** History of the Malay language, the Malay language from the 4th to the 14th century ** Indonesi ...
,
Sundanese Sundanese may refer to: * Sundanese people * Sundanese language * Sundanese script Standard Sundanese script (''Aksara Sunda Baku'', ) is a writing system which is used by the Sundanese people. It is built based on Old Sundanese script (' ...
, Tagalog,
Thai Thai or THAI may refer to: * Of or from Thailand, a country in Southeast Asia ** Thai people, the dominant ethnic group of Thailand ** Thai language, a Tai-Kadai language spoken mainly in and around Thailand *** Thai script *** Thai (Unicode block ...
, or Javanese), as well as the family name ''Gekkonidae'' and the generic term ''gecko'' come from. Most of the time, the call is often preceded by a quick "cackling", similar to the chirping sounds made by
house gecko __NOTOC__ ''Hemidactylus'' is a genus of the common gecko family, Gekkonidae. It has 189 described species, newfound ones being described every few years. These geckos are found in all the tropical regions of the world, extending into the subtrop ...
s albeit much lower in pitch. When threatened or alarmed, tokay geckos usually "bark" while opening their mouth in a defensive posture. The tokay gecko's call is also responsible for the name given to it by Filipino residents: "Tuko," and by U.S. soldiers during the
Vietnam War The Vietnam War (also known by #Names, other names) was a conflict in Vietnam, Laos, and Cambodia from 1 November 1955 to the fall of Saigon on 30 April 1975. It was the second of the Indochina Wars and was officially fought between North Vie ...
, the "Fuck-you lizard". Light and temperature can affect its vocalizations. The most frequent calling occurs in May at dusk, and the second peak of call frequency occurs in May at dawn. Vocalizations and associated behavior were strongly affected by ambient temperature in both the lab and field and could thus play a role in regulating animal energetic metabolism


Conservation and relationship with humans

The tokay gecko is culturally significant in many East Asian countries. Regional folklore has attributed supernatural powers to the gecko. In
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
it is a symbol of good luck and fertility. It is believed to be descended from
dragon A dragon is a reptilian legendary creature that appears in the folklore of many cultures worldwide. Beliefs about dragons vary considerably through regions, but dragons in western cultures since the High Middle Ages have often been depicted as ...
s.Naish, D
People are modifying monitors to make gargantuan geckos.
''Scientific American'' Blog 16 April 2015.
This species is poached for the medicinal trades in parts of Asia. The tokay gecko is an ingredient in
traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of action ...
, known as ''Ge Jie'' (蛤蚧). It is believed to nourish the kidneys and lungs, beliefs that are not substantiated by medical science. The animal remains highly sought after in China, Hong Kong, Taiwan, Vietnam, Malaysia, Singapore and other parts of Asia with Chinese communities, to the point where unscrupulous merchants have taken to disfiguring
monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and one species is also found in the Americas as an invasive species. About 80 species are recogn ...
s with prosthetics to pass them off as colossal tokay gecko specimens. From 2009 to 2011, the poaching of tokay geckos intensified because of a short-lived belief that they were an effective
HIV The human immunodeficiency viruses (HIV) are two species of ''Lentivirus'' (a subgroup of retrovirus) that infect humans. Over time, they cause acquired immunodeficiency syndrome (AIDS), a condition in which progressive failure of the immune ...
cure. The tokay gecko is quickly becoming a threatened species in the Philippines because of indiscriminate hunting. Collecting, transporting and trading in geckos without a license can be punishable by up to 12 years in jail and a fine of up to
The Philippine peso sign (₱) is the currency symbol used for the Philippine peso, the official currency of the Philippines. The symbol resembles a Roman letter P with two horizontal strokes. It differs from the currency symbol used for the pe ...
1 million under Republic Act 9147, in addition to other applicable international laws. However, the trade runs unchecked because of the sheer number of illegal traders and reports of lucrative deals.
Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of va ...
buyers and other foreign nationals are rumored to pay thousands of dollars for large specimens, because of their alleged medicinal value or as commodities in the illegal wildlife trade. The species is protected under Appendix II of the
Convention on International Trade in Endangered Species CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
(CITES) meaning international trade (including in parts and derivatives) is subject to the CITES permitting system.


Captivity

Tokay geckos are becoming more popular as pets because of their striking colors and large size. Most of them are wild-caught imports, but captive-bred ones are becoming more common. Wild-caught adults can be difficult to keep because of their aggressive nature and powerful bite, but captive-bred juveniles can be less aggressive if handled from a young age. When well cared for, tokay geckos can live up to 15–20 years.


References


External links



Reptile Parkreptilepark.com.au.

EcologyAsia.
Tokay Gecko Care.
TokayGecko.org
The sound of a Balinese geko in Sanur. Recorded October 2004.
SoundCloud.
Nonnatives - Tokay Gecko.
Florida Fish and Wildlife Conservation Commission.

Australian Reptile Park. {{Taxonbar, from=Q331213 Geckos Gekko Reptiles of Southeast Asia Reptiles of Bangladesh Reptiles of China Reptiles of Taiwan Reptiles described in 1758 Taxa named by Carl Linnaeus