Garrya Congdonii
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''Garrya congdonii'', the chaparral silktassel or Congdon silktassel, a fairly common
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
native to the northern
California Coast Ranges The Coast Ranges of California span from Del Norte or Humboldt County, California, south to Santa Barbara County. The other three coastal California mountain ranges are the Transverse Ranges, Peninsular Ranges and the Klamath Mountains. P ...
, is one of a small biological family of approximately twenty known species in the family ''
Garryaceae Garryaceae is a small family of plants known commonly as the silktassels.Garryaceae.
Flora o ...
'', most of which are Garrya. While the female and male
sexual organ A sex organ (or reproductive organ) is any part of an animal or plant that is involved in sexual reproduction. The reproductive organs together constitute the reproductive system. In animals, the testis in the male, and the ovary in the female, a ...
s of Congdon silktassel are on separate plants, the pendant male
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s are much more showy. This plant is reasonably attractive and neat enough in its growing habit to be appealing as a
landscape A landscape is the visible features of an area of land, its landforms, and how they integrate with natural or man-made features, often considered in terms of their aesthetic appeal.''New Oxford American Dictionary''. A landscape includes the ...
species. It is stocked commonly at commercial plant nurseries. All Garrya are associated with warm temperate regions of North America.


Description

240px, Leaves and spent blooms Congdon silktassel has a multi-furcate trunk branching structure yielding an almost spherical form. The height can attain four meters, but more commonly averages two to three meters in the wild. Congdon silktassel, as all the genus Garrya, have opposite leaves that have a tough leathery feel, glossy green on top, but paler, slightly furry and duller on the underside. The
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
flowers are concentrated in
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s which cascade downward as
ament Ament may refer to: * Catkin, a cylindrical flower cluster * Ament (surname) * Imentet Imentet (Ament, Amentet or Imentit, meaning "She of the West") was a goddess in ancient Egyptian religion representing the necropolises west of the Nile. ...
s of approximately three to five centimeters in length. While the Congdon silktassel manifests separate male and female plants, the pendant male catkins are much more evident, up to 25 centimeters long; those of the female are shorter and silver-grey. Although the
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s bloom in late January and February, dried
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s remain on the tree well into summer as light gray decorations. The plant has smooth dark bark, dark-greenish when young, but with age the bark roughens. New
twig A twig is a thin, often short, branch of a tree or bush. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the twig bark are ...
s are green and moderately stout. For
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
late flowers, above each small bract there is a solitary flower inside the inflorescence. This plant produces tiny dark
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s that can become airborne as small fluffy wind-blown cotton. The ripened purple to black
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
of about seven millimeters in diameter has a hard desiccated shell, but is somewhat fleshy on the interior. In the case of
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
ate inflorescences, there are a total of four stamens per flower; moreover, above each bract pair there is a triplet of flowers. The unique characteristics of Congdon silktassel are its leathery
convex Convex or convexity may refer to: Science and technology * Convex lens, in optics Mathematics * Convex set, containing the whole line segment that joins points ** Convex polygon, a polygon which encloses a convex set of points ** Convex polytope ...
green (with a very slight yellowish tinge) leaves with wavy leaf margins, coupled with dense individual hairs on the leaf undersides that are scarcely distinguishable with a hand lens. Its leaf blades are six to eight centimeters in length, and has petioles which range in length from six to twelve millimeters. For identification purposes, Congdon silktassel has leaf hairs that are distinguishable with a hand lens, and both leaf blades and petioles are about two thirds the size of those on Coast silktassel.


Habitat

Congdon silktassel is found in several plant communities, principally in drier inner coastal California ranges no more than 20 miles from the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
; the main associations are Coastal
chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...
, Mixed
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zone ...
(near a chaparral area) and Northern coastal
sage Sage or SAGE may refer to: Plants * ''Salvia officinalis'', common sage, a small evergreen subshrub used as a culinary herb ** Lamiaceae, a family of flowering plants commonly known as the mint or deadnettle or sage family ** ''Salvia'', a large ...
scrub. This plant can tolerate moderately heavy
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay par ...
soils and serpentine areas, but likes soils pH to lie in the range of six to eight. An example of serpentine soils habitat is in The Cedars of Sonoma County, California.C.Michael Hogan. 2010
''Leather oak''. eds. mark McGinley & C.J.Cleveland. Encyclopedia of Earth. National Council for Science and the Environment. Washington DC
/ref> This plant is rather resistant to being grazed by deer or rabbits, and is hardy to cold temperatures of about 15 degrees Fahrenheit. Congdon silktassel is moderately drought-tolerant, but is more luxuriant with rainfall of about 22 inches per annum.


Differences with other ''Garrya'' species

Congdon silktassel is usually found at elevations above 200 meters. This species is usually found within the mountains of the inner
Pacific Coast Pacific coast may be used to reference any coastline that borders the Pacific Ocean. Geography Americas Countries on the western side of the Americas have a Pacific coast as their western or southwestern border, except for Panama, where the Pac ...
range, such as Mount Hamilton ranging north to the inner coast range in
Sonoma County Sonoma County () is a county (United States), county located in the U.S. state of California. As of the 2020 United States Census, its population was 488,863. Its county seat and largest city is Santa Rosa, California, Santa Rosa. It is to the n ...
, which is where the specimen photographed herein was found. Coast silktassel (''Garrya elliptica'') is most closely related species. Coast silktassel has similar wavy undulated entire leaf margins with somewhat darker glossier leaves, and the leaf topsides are more waxy and not quite so leathery as Congdon. The whitish hairs on the leaf underside of Coast silktassel are not readily visible with a hand lens as for Congdon silktassel. Coast silktassel is most commonly seen growing as individual plants rather than in groves, and can be found in the
Montara Mountain Montara Mountain, positioned between the unincorporated community of Montara, California, to the southwest and the city of Pacifica, California, to the north, forms the northern spur of the Santa Cruz Mountains, a narrow mountain range running the ...
range, San Bruno Mountain and outer coastal ranges in
Marin County Marin County is a county located in the northwestern part of the San Francisco Bay Area of the U.S. state of California. As of the 2020 census, the population was 262,231. Its county seat and largest city is San Rafael. Marin County is acros ...
such as
Mount Tamalpais Mount Tamalpais (; ; Miwok languages, Miwok: ''Támal Pájiṣ''), known locally as Mount Tam, is a mountain, peak in Marin County, California, Marin County, California, United States, often considered symbolic of Marin County. Much of Mount Tama ...
. Both Fremont silktassel (''Garrya fremontii'') and Ashy silktassel (''Garrya flavescens'') have similar fruit characteristics, but have a flat leaf margin. Fremont silktassel has yellow-green leaves and fruits that are almost lacking in hairs. Fremont silktassel is found from the Mount Hamilton area and
Loma Prieta Loma Prieta (from Spanish ''loma'' -hill, ''prieta'' -dark) is high and is the highest peak in the Santa Cruz Mountains in Northern California. Although the 1989 Loma Prieta earthquake was named for this mountain, the actual epicenter was fi ...
area ranging north to Marin County (for example, on Mount Tamalpais. Ashy silktassel (''Garrya flavescens pallida'') has grayish leaves and is reported to occur in the
Alameda County Alameda County ( ) is a List of counties in California, county located in the U.S. state of California. As of the 2020 United States Census, 2020 census, the population was 1,682,353, making it the 7th-most populous county in the state and List ...
hills.


See also

*
Joseph Whipple Congdon Joseph Whipple Congdon (April 13, 1834 – April 5, 1910) was a lawyer by trade who contributed significantly to early botanical exploration in California, particularly in the Yosemite region, where he resided in Mariposa County, California, Marip ...


References

{{Taxonbar, from=Q5524256 Garryales Endemic flora of California Natural history of the California Coast Ranges Natural history of the California chaparral and woodlands Natural history of the San Francisco Bay Area Garden plants of North America Drought-tolerant plants Flora without expected TNC conservation status