Galve, Teruel
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Galve is a municipality located in the
province of Teruel Teruel (Catalan: ''Terol'' ) is a province of Aragon, in the northeast of Spain. The capital is Teruel. It is bordered by the provinces of Tarragona, Castellón, Valencia (including its exclave Rincón de Ademuz), Cuenca, Guadalajara, and Zar ...
,
Aragon Aragon ( , ; Spanish and an, Aragón ; ca, Aragó ) is an autonomous community in Spain, coextensive with the medieval Kingdom of Aragon. In northeastern Spain, the Aragonese autonomous community comprises three provinces (from north to sou ...
, Spain. According to the 2006
census A census is the procedure of systematically acquiring, recording and calculating information about the members of a given population. This term is used mostly in connection with national population and housing censuses; other common censuses incl ...
(
INE INE, Ine or ine may refer to: Institutions * Institut für Nukleare Entsorgung, a German nuclear research center * Instituto Nacional de Estadística (disambiguation) * Instituto Nacional de Estatística (disambiguation) * Instituto Nacional Elec ...
), the municipality has a population of 145 inhabitants. There is an important paleontological site.


Paleontological significance

The Galve area is a productive area full of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
sites well known for its faunal assemblage of dinosaurs, notably the
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
s and
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. This region is also noted for its
lissamphibia The Lissamphibia is a group of tetrapods that includes all modern amphibians. Lissamphibians consist of three living groups: the Salientia (frogs, toads, and their extinct relatives), the Caudata (salamanders, newts, and their extinct relatives), ...
ns,
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s and
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...
s, as well as diverse
multituberculate Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, a ...
s,
dryolestoid Dryolestida is an extinct order of mammals, primarily and possibly exclusively known from the Jurassic and Cretaceous. They are considered members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they d ...
s, and
spalacotheriid Spalacotheriidae is a family of extinct mammals belonging to the paraphyletic group ' Symmetrodonta'. They lasted from the Early Cretaceous to the Campanian in North America, Europe, Asia and North Africa. Spalacotheriids are characterised by ha ...
s. Spanish paleontologists have worked this site for years, including José Ignacio Canudo, José Luis Barco, Gloria Cuenca-Bescós, and José Ignacio Ruiz-Omeñaca. The Galve region sediments date back to the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
, with five formations ranging in age from
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
to
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
listed below: * Higueruelas Formation (
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
) *
Villar del Arzobispo Formation The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Middl ...
(upper
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
-middle
Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...
) *
El Castellar Formation The El Castellar Formation is a geological formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recover ...
(upper
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
) *
Camarillas Formation The Camarillas Formation is a geological formation in the Teruel Province of Aragón, Spain whose strata date back to the Early Cretaceous (Barremian stage). The sandstones, mudstones and conglomerates of the formation, that due to syn-sediment ...
(lower
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
) *
Artoles Formation The Artoles Formation is a geological formation in Teruel and La Rioja, Spain whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel et al., 2004, p.561 Vert ...
(upper
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
-lower
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
)


Fauna


Dinosaurs

Theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
s ( spinosaurines),
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
s, and
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
ns have been reported from the Galve region.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
belonging to
baryonychine Baryonychinae is an extinct clade or subfamily of spinosaurids from the Early Cretaceous (Valanginian-Albian) of Britain, Portugal, and Niger. In 2021, it consisted of six genera: '' Ceratosuchops'', ''Cristatusaurus'', '' Riparovenator'', '' ...
s,
allosauroid Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among th ...
s,
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
s and indeterminable
coelurosaur Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrann ...
s have also been recovered at some of the Galve sites. Those indeterminable coelurosaur teeth are from the Galve outcrops of the
El Castellar Formation The El Castellar Formation is a geological formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recover ...
, and bear some
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n resemblance.


=Sauropods

= Sauropods from the Galve region include '' Aragosaurus ischiaticus'', '' Galveosaurus herreroi'', and ''
Turiasaurus riodevensis ''Turiasaurus'' (meaning " Turia lizard") is a genus of sauropod dinosaurs. It is known from a single fossil specimen representing the species ''Turiasaurus riodevensis'', found in the Kimmeridgian Villar del Arzobispo Formation of Teruel, Spain ...
''. Isolated remains have also been referred to
camarasaurid Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thic ...
s,
brachiosaurid The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the le ...
s, and
diplodocid Diplodocids, or members of the family Diplodocidae ("double beams"), are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including ''Diplodocus'' and ''Supersaurus'', some of which may have ...
s. ''Aragosaurus'' has been identified as a
camarasaurid Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thic ...
, a
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
, and recently, a
eusauropod Eusauropoda (meaning "true sauropods") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of ''Shunosaurus'', and possibly ''Barapasaurus ...
of uncertain affinities. ''Galveosaurus'' was originally described as a cetiosaurid but later argued to be a basal
macronaria Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ...
n, and ''Turiasaurus'' was argued by its describers to represent a new
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
close to neosauropod ancestry,
Turiasauria Turiasauria is an unranked clade of basal sauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa. Description Turiasauria was originally erected by Royo-Torres et al. (2006) to include '' ...
. A new study also concluded that ''Galveosaurus'' was a turiasaur, and not a cetiosaurid or macronarian. ''Galveosaurus'' was published in
2005 File:2005 Events Collage V2.png, From top left, clockwise: Hurricane Katrina in the Gulf of Mexico; the Funeral of Pope John Paul II is held in Vatican City; "Me at the zoo", the first video ever to be uploaded to YouTube; Eris was discovered in ...
by Barbara, naming it ''Galveosaurus herreroi'', in the
journal A journal, from the Old French ''journal'' (meaning "daily"), may refer to: *Bullet journal, a method of personal organization *Diary, a record of what happened over the course of a day or other period *Daybook, also known as a general journal, a ...
Zootaxa.Sánchez-Hernández, B. 2005. ''Galveosaurus herreroi'', a new sauropod dinosaur from Villar del Arzobispo Formation (Tithonian-Berriasian) of Spain. ''Zootaxa'' 1034, 1-20. Late that same year, a team led by José Luis Barco published a less formal article in the magazine ''Naturaleza Aragonesa'' (Barco et al. 2005), and here they named the same taxon ''Galvesaurus herreroi''(a difference of one letter). A ''
Camarasaurus ''Camarasaurus'' ( ) was a genus of quadrupedal, herbivorous dinosaurs and is the most common North American sauropod fossil. Its fossil remains have been found in the Morrison Formation, dating to the Late Jurassic epoch (Kimmeridgian to Titho ...
''-like taxon is represented only by teeth from the
El Castellar Formation The El Castellar Formation is a geological formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recover ...
. These teeth bear a strong resemblance to the teeth of ''Camarasaurus'' than the teeth of any other sauropod. However, their younger age (
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
-
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
) makes them substantially younger than ''Camarasaurus'', and hence likely to belong to a separate
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
.


=Heterodontosaurids

=
Heterodontosaurid Heterodontosauridae is a family of ornithischian dinosaurs that were likely among the most basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly found to be primitive, outside of the group ...
s have also been attributed to the Galve region. Heterodontosaurids are a mostly
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
, best known from
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number of ...
.


Crocodilians

Numerous
crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
ns have been found in Galve. One particularly notable find is an outcrop of the
Villar del Arzobispo Formation The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Middl ...
at El Cantalar that preserves a trackway produced by a
crocodyliform Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
that measures approximately 12 meters in length, comparable in size to ''
Deinosuchus ''Deinosuchus'' () is an extinct genus of alligatoroid crocodilian, related to modern alligators and caimans, that lived 82 to 73 million years ago (Ma), during the late Cretaceous period. The name translates as "terrible crocodile" and is ...
'', ''
Sarcosuchus ''Sarcosuchus'' (; ) is an extinct genus of crocodyliform and distant relative of living crocodilians that lived during the Early Cretaceous, from the late Hauterivian to the early Albian, 133 to 112 million years ago of what is now Africa and ...
'', and the unique ''
Stomatosuchus ''Stomatosuchus'' (type species ''S. inermis'') is an extinct stomatosuchid neosuchian from the Late Cretaceous ( Cenomanian) of Egypt. Much of what is known about ''Stomatosuchus'' has been inferred from the related genus ''Laganosuchus''. De ...
''. Furthermore,
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
,
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s, and
bone A bone is a Stiffness, rigid Organ (biology), organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red blood cell, red and white blood cells, store minerals, provid ...
s belonging to
atoposaurid Atoposauridae is a Family (biology), family of crocodile-line archosaurs belonging to Neosuchia. The majority of the family are known from Late Jurassic to Early Cretaceous marine deposits in France, Portugal, and Bavaria in southern Germany. The ...
s, ''
Bernissartia ''Bernissartia'' ('of Bernissart') is an extinct genus of neosuchian crocodyliform that lived in the Early Cretaceous, around 130 million years ago. At only in length, ''Bernissartia'' is one of the smallest crocodyliforms that ever lived. It ...
'', and possibly pholidosaurids and teleosaurids are all known from the Galve region, illustrating the amount of crocodilian material preserved at the site.Sánchez-Hernández, B., Benton, M. J. & Naish, D.
2007 File:2007 Events Collage.png, From top left, clockwise: Steve Jobs unveils Apple's first iPhone; TAM Airlines Flight 3054 overruns a runway and crashes into a gas station, killing almost 200 people; Former Pakistani Prime Minister of Pakistan, Pr ...
. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. ''Palaeogeography, Palaeoclimatology, Palaeoecology'' 249, 180-215.


Pterosaurs

Pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
teeth have also been uncovered in the Galve region, although more substantial remains are rare. However, there are enough remains at the sites to determine that the Galve region hosted a wide variety of pterosaurs at one point. Pterosaurs that have been discovered at this site include
ornithocheirid Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cr ...
s,
dsungaripterid Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings. Classification In 1964 Young created a family to place the recentl ...
s, possible gnathosaurines, and
istiodactylid Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus '' Ornithodesmus''. Systematics and distribution Remains of taxa that can be c ...
s.


References

{{authority control Municipalities in the Province of Teruel Mesozoic paleontological sites of Europe Geography of Spain