Galve, Spain
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Galve is a municipality located in the
province of Teruel Teruel ( Catalan: ''Terol'' ) is a province of Aragon, in the northeast of Spain. The capital is Teruel. It is bordered by the provinces of Tarragona, Castellón, Valencia (including its exclave Rincón de Ademuz), Cuenca, Guadalajara, and Z ...
,
Aragon Aragon ( , ; Spanish and ; ) is an autonomous communities of Spain, autonomous community in Spain, coextensive with the medieval Kingdom of Aragon. In northeastern Spain, the Aragonese autonomous community comprises three provinces of Spain, ...
, Spain. According to the 2006
census A census (from Latin ''censere'', 'to assess') is the procedure of systematically acquiring, recording, and calculating population information about the members of a given Statistical population, population, usually displayed in the form of stati ...
( INE), the municipality has a population of 145 inhabitants. There is an important paleontological site.


Paleontological significance

The Galve area is a productive area full of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
sites well known for its faunal assemblage of dinosaurs, notably the
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
s and
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s. This region is also noted for its
lissamphibia The Lissamphibia (from Greek λισσός (lissós, "smooth") + ἀμφίβια (amphíbia), meaning "smooth amphibians") is a group of tetrapods that includes all modern amphibians. Lissamphibians consist of three living groups: the Salientia ( ...
ns,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s and
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s, as well as diverse
multituberculate Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, a ...
s,
dryolestoid Dryolestida is an extinct order of mammals, known from the Jurassic and Cretaceous. They are considered Basal (phylogenetics), basal members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they develop ...
s, and
spalacotheriid Spalacotheriidae is a family of extinct mammals belonging to the paraphyletic group 'Symmetrodonta'. They lasted from the Early Cretaceous to the Campanian in North America, Europe, Asia and North Africa. Spalacotheriids are characterised by hav ...
s. Spanish paleontologists have worked this site for years, including
José Ignacio Canudo José is a predominantly Spanish and Portuguese form of the given name Joseph. While spelled alike, this name is pronounced very differently in each of the two languages: Spanish ; Portuguese (or ). In French, the name ''José'', pronounced , ...
,
José Luis Barco José is a predominantly Spanish and Portuguese language, Portuguese form of the given name Joseph. While spelled alike, this name is pronounced very differently in each of the two languages: Spanish ; Portuguese (or ). In French, the name ''J ...
,
Gloria Cuenca-Bescós Gloria may refer to: Arts and entertainment Music Christian liturgy and music * Gloria in excelsis Deo, the Greater Doxology, a hymn of praise * Gloria Patri, the Lesser Doxology, a short hymn of praise ** Gloria (Handel) ** Gloria (Jenkins ...
, and
José Ignacio Ruiz-Omeñaca José is a predominantly Spanish and Portuguese form of the given name Joseph. While spelled alike, this name is pronounced very differently in each of the two languages: Spanish ; Portuguese (or ). In French, the name ''José'', pronounced , ...
. The Galve region sediments date back to the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
, with five formations ranging in age from
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
to
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
listed below: * Higueruelas Formation (
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
) *
Villar del Arzobispo Formation The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Mid ...
(upper
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
-middle
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
) *
El Castellar Formation The El Castellar Formation is a geological Formation (geology), formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils ...
(upper
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
) *
Camarillas Formation The Camarillas Formation is a geological formation in the Teruel Province of Aragón, Spain whose strata date back to the Early Cretaceous (Barremian stage). The sandstones, mudstones and conglomerates of the formation, that due to syn-sedimen ...
(lower
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) * Artoles Formation (upper
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
-lower
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
)


Fauna


Dinosaurs

Theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
s (
spinosaurine Spinosauridae (or spinosaurids) is a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. Their remains have ...
s),
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
s, and
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
ns have been reported from the Galve region.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
belonging to
baryonychine Baryonychinae is an extinct clade or subfamily of spinosaurids from the Early Cretaceous of Europe and West Africa. The clade was named by Charig & Milner in 1986 and defined by Sereno et al. in 1998 and Holtz et al. in 2004 as all taxa more close ...
s,
allosauroid Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods. While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
s,
dromaeosaurid Dromaeosauridae () is a family (biology), family of feathered coelurosaurian Theropoda, theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous period (geology), Period. The name Drom ...
s and indeterminable
coelurosaur Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
s have also been recovered at some of the Galve sites. Those indeterminable coelurosaur teeth are from the Galve outcrops of the
El Castellar Formation The El Castellar Formation is a geological Formation (geology), formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils ...
, and bear some
avia AVIA () is a Soviet/Russian experimental pop band formed in Leningrad in 1986. AVIA released four studio albums and led the first wave of the Soviet bands which made their breakthrough in the West in the late 1980s. Band history AVIA was formed ...
n resemblance.


Sauropods

Sauropods from the Galve region include '' Aragosaurus ischiaticus'', '' Galveosaurus herreroi'', and ''
Turiasaurus riodevensis ''Turiasaurus'' (meaning " Turia lizard") is a genus of sauropod dinosaurs. It is known from a single fossil specimen representing the species ''Turiasaurus riodevensis'', found in the Kimmeridgian Villar del Arzobispo Formation of Teruel, Spain ...
''. Isolated remains have also been referred to camarasaurids,
brachiosaurid The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the l ...
s, and
diplodocid Diplodocids, or members of the family Diplodocidae ("double beams"), are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including '' Diplodocus'' and '' Supersaurus'', some of which may ha ...
s. ''Aragosaurus'' has been identified as a camarasaurid, a
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
, and recently, a
eusauropod Eusauropoda (meaning "True Lizard Foot") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of ''Shunosaurus'', and possibly ''Barapasaur ...
of uncertain affinities. ''Galveosaurus'' was originally described as a cetiosaurid but later argued to be a basal
macronaria Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ...
n, and ''Turiasaurus'' was argued by its describers to represent a new
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
close to neosauropod ancestry,
Turiasauria Turiasauria is an unranked clade of eusauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa. Description Turiasauria was originally erected by Royo-Torres et al. (2006) to include '' Tur ...
. A new study also concluded that ''Galveosaurus'' was a turiasaur, and not a cetiosaurid or macronarian. ''Galveosaurus'' was published in
2005 2005 was designated as the International Year for Sport and Physical Education and the International Year of Microcredit. The beginning of 2005 also marked the end of the International Decade of the World's Indigenous Peoples, Internationa ...
by Barbara, naming it ''Galveosaurus herreroi'', in the
journal A journal, from the Old French ''journal'' (meaning "daily"), may refer to: *Bullet journal, a method of personal organization *Diary, a record of personal secretive thoughts and as open book to personal therapy or used to feel connected to onesel ...
Zootaxa.Sánchez-Hernández, B. 2005. ''Galveosaurus herreroi'', a new sauropod dinosaur from Villar del Arzobispo Formation (Tithonian-Berriasian) of Spain. ''Zootaxa'' 1034, 1-20. Late that same year, a team led by
José Luis Barco José is a predominantly Spanish and Portuguese language, Portuguese form of the given name Joseph. While spelled alike, this name is pronounced very differently in each of the two languages: Spanish ; Portuguese (or ). In French, the name ''J ...
published a less formal article in the magazine ''Naturaleza Aragonesa'' (Barco et al. 2005), and here they named the same taxon ''Galvesaurus herreroi''(a difference of one letter). A ''
Camarasaurus ''Camarasaurus'' ( ) is a genus of sauropod dinosaur that lived in North America during the Late Jurassic period. Its fossil remains have been found in the Morrison Formation, dating to the Kimmeridgian and Tithonian ages of the Jurassic, betwe ...
''-like taxon is represented only by teeth from the
El Castellar Formation The El Castellar Formation is a geological Formation (geology), formation in La Rioja and Teruel, Spain whose strata date back to the possibly the Valanginian to the Barremian stages of the Early Cretaceous. Dinosaur remains are among the fossils ...
. These teeth bear a strong resemblance to the teeth of ''Camarasaurus'' than the teeth of any other sauropod. However, their younger age (
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
-
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) makes them substantially younger than ''Camarasaurus'', and hence likely to belong to a separate
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
.


=Heterodontosaurids

= Heterodontosaurids have also been attributed to the Galve region. Heterodontosaurids are a mostly
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic ...
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, best known from
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
.


Crocodilians

Numerous
crocodilia Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...
ns have been found in Galve. One particularly notable find is an outcrop of the
Villar del Arzobispo Formation The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Mid ...
at El Cantalar that preserves a trackway produced by a
crocodyliform Crocodyliformes is a clade of Crurotarsi, crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the ...
that measures approximately 12 meters in length, comparable in size to ''
Deinosuchus ''Deinosuchus'' is an extinct genus of eusuchian, either an Alligatoroidea, alligatoroid Crocodilia, crocodilian or a stem-group crocodilian, which lived during the Late Cretaceous around . The first remains were discovered in North Carolina ...
'', ''
Sarcosuchus ''Sarcosuchus'' (), from Ancient Greek σάρξ (''sárx''), meaning "flesh", and σούχος (''soúkhos''), meaning "crocodile", is an extinct genus of crocodyliform a that lived during the Early Cretaceous, from the late Hauterivian to the ...
'', and the unique ''
Stomatosuchus ''Stomatosuchus'' (meaning "mouth crocodile") is an extinct stomatosuchid neosuchian from the Late Cretaceous (Cenomanian) Bahariya Formation of Egypt. The type and only species is ''S. inermis''. Much of what is known about ''Stomatosuchus'' ha ...
''. Furthermore,
teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
,
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s, and
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
s belonging to atoposaurids, ''
Bernissartia ''Bernissartia'' ('of Bernissart') is an extinct genus of neosuchian crocodyliform that lived in the Early Cretaceous, around 130 million years ago. At only in length, ''Bernissartia'' is one of the smallest crocodyliforms that ever lived. It ...
'', and possibly
pholidosaurid Pholidosauridae is an extinct Family (biology), family of aquatic neosuchian mesoeucrocodylian crocodylomorphs. Fossils have been found in Europe (Denmark, England, France, Germany, Spain and Sweden), Africa (Algeria, Niger, Mali, Morocco and Tu ...
s and teleosaurids are all known from the Galve region, illustrating the amount of crocodilian material preserved at the site.Sánchez-Hernández, B., Benton, M. J. & Naish, D.
2007 2007 was designated as the International Heliophysical Year and the International Polar Year. Events January * January 1 **Bulgaria and Romania 2007 enlargement of the European Union, join the European Union, while Slovenia joins the Eur ...
. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. ''Palaeogeography, Palaeoclimatology, Palaeoecology'' 249, 180–215.


Pterosaurs

Pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
teeth have also been uncovered in the Galve region, although more substantial remains are rare. However, there are enough remains at the sites to determine that the Galve region hosted a wide variety of pterosaurs at one point. Pterosaurs that have been discovered at this site include
ornithocheirid Anhangueridae (alternatively called Ornithocheiridae, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the ...
s,
dsungaripterid Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings, and were commonly interpreted as durophagous and possibly piscivorou ...
s, possible
gnathosaurine Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early ...
s, and
istiodactylid Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus '' Ornithodesmus''. Systematics and distribution Remains of taxa that can be c ...
s.


See also

*
List of municipalities in Teruel This is a list of the municipalities in the province of Teruel in the autonomous community of Aragon, Spain. There are 236 municipalities in the province. List See also * Geography of Spain * List of cities in Spain * List of Aragonese comarcas ...


References

{{authority control Municipalities in the Province of Teruel Mesozoic paleontological sites of Europe Geography of Spain