Furnarius rufus
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The rufous hornero (''Furnarius rufus'') is a medium-sized ovenbird in the family Furnariidae. It occurs in eastern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, and is the
national bird This is a list of national birds, including official birds of overseas territories and other states described as nations. Most species in the list are officially designated. Some species hold only an "unofficial" status. National birds See al ...
of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. Also known as the red ovenbird, it is common in savannas, second-growth scrub, pastures and agricultural land and is synanthropic. Its range includes midwestern, southeastern and southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
and northern and central
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, extending as far south as northern Patagonia. The species is most closely related to the crested hornero of Paraguay and Argentina. There are four accepted
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. The rufous hornero is medium-sized with a square tail and very slightly decurved bill. The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is overall reddish brown with a dull brown crown and a whitish throat. Sexes are alike and juvenile birds are slightly paler below (probably because they are cleaner). Rufous horneros feed on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s and other
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s obtained by foraging on the ground while walking. They sometimes feed on scraps such as bread crumbs. Songs in the rufous hornero are sexually distinct. The rapid trill that is usually heard as part of the duet is faster in the male, slower in the female, and both beat their wings at their sides while singing and the wings beat at the same rate as their trill. Thus, while watching an observer may identify the sex by how fast their wings beat while singing.


Predators

Predators of adult and young rufous horneros include birds of prey such as the black-chested buzzard-eagle ''
Buteo melanoleucus The black-chested buzzard-eagle (''Geranoaetus melanoleucus'') is a bird of prey of the hawk and eagle family (Accipitridae). It lives in open regions of South America. This species is also known as the black buzzard-eagle, the gray buzzard-ea ...
'', small mammals, domestic cats, and a number of species of snakes and possibly lizards.Cf. Wikiaves photo, taken in Brasilia

/ref> However, its covered nest probably reduces predation risk.


Breeding

The rufous hornero breeds in the
austral summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, with ...
, laying eggs between August and December, raising nestlings soon after, and the young may stay in their natal territory until the following breeding season. The species is
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
and the pair bond is long term, sometimes for life. The nest of the species is typical for the genus, a large thick clay "oven" placed on a tree, or man made structures such as fenceposts, telephone poles or buildings. Pairs remain together throughout the year and will work on the nest during that time; nests can be constructed in as little as 5 days but usually take longer, occasionally months, to complete. A clutch generally contains two to four eggs. The eggs are laid every second day and incubated for 14–18 days. Chicks are fed for 23–26 days before
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
; young birds remain in the parental territory for around 6 months after fledging and sometimes until the following breeding season. Both parents incubate eggs and feed the young. Horneros may or may not reuse nests, therefore it is quite common to see several nests close to each other (or even atop older nests) at the same nesting site. However, a formerly unused nest may be repaired for a new breeding season.


Status

The rufous hornero has benefited from human changes to the environment and many live in highly modified habitat, such as city suburbs. In turn abandoned hornero nests may be of benefit to various other species of birds that nest in its unused "ovens". The
saffron finch The saffron finch (''Sicalis flaveola'') is a tanager from South America that is common in open and semi-open areas in lowlands outside the Amazon Basin. They have a wide distribution in Colombia, northern Venezuela (where it is called "canar ...
is one species that commonly nests in old ovenbird nests. The rufous hornero is a familiar sight over much of its range and has been adopted as the
national bird This is a list of national birds, including official birds of overseas territories and other states described as nations. Most species in the list are officially designated. Some species hold only an "unofficial" status. National birds See al ...
of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
and
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
. It is not threatened by human activities and is listed as
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. Rufous hornero (Red ovenbird)(Furnarius rufus) and nest.JPG, building nest
The Pantanal,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
Rufous hornero (Red ovenbird)(Furnarius rufus) and nest (2).JPG, outside completed nest
The Pantanal,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...


Taxonomy and systematics

The first notes taken on the species were made by
Philibert Commerson Philibert Commerson (; 18 November 1727 – 14 March 1773), sometimes spelled Commerçon by contemporaries, was a French naturalist, best known for accompanying Louis Antoine de Bougainville on his voyage of circumnavigation in 1766–1769. ...
in 1767, from a specimen obtained at
Barragán cove The Ensenada de Barragán ("Barragán's Bay, or Cove") is a small bay on the Río de la Plata, some southeast of Buenos Aires, Argentina. The area is in the Ensenada district of Buenos Aires province, centred on the city of the Ensenada. The ol ...
during Louis Antoine de Bougainville's expedition. Commerson named the bird as ''Turdus fulvus'' and his notes were later published by
Georges Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent Fr ...
in his '' Histoire Naturelle'' in 1779. However, the rufous hornero was first scientifically described, as ''Merops rufus'', by the German naturalist
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...
in the 13th edition of ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' published in 1788. In 1816, Louis Jean Pierre Vieillot established the genus '' Furnarius'' in his ''Analyse d'une nouvelle ornithologie élémentaire'' and included the rufous hornero on it, although Vieillot did not directly rename the rufous hornero as ''Furnarius rufus''. Its current
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
was used for the first time in ornithology by John Gould in his '' Zoology of the Voyage of H.M.S. Beagle'' in 1841. Nowadays the rufous hornero integrates the genus ''Furnarius'' with other five species. They are all native to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
and build mud nests that resemble old wood-fired ovens. Its closest relative is the crested hornero, which is considered as its sister species due to similar behavior and plumage pattern. The derivation of the current genus name, ''Furnarius'', is from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''furnus'', meaning "an oven". The Spanish word "hornero" similarly comes from horno, meaning "oven". Its
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
comes from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''rufum'', meaning "red" or "reddish". It is also known as the red ovenbird.


Subspecies

Five subspecies of the rufous hornero are generally recognized,«Ovenbirds & woodcreepers»
IOC World Bird List Version 5.1. Retrieved 22 March 2015.
based upon plumage and size: *''F. r. rufus'', the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, described by
Gmelin Gmelin may refer to: * Gmelin's test, a chemical test * Gmelin database, a German handbook/encyclopedia of inorganic compounds initiated by Leopold Gmelin People * Carl Christian Gmelin (1762–1837), German botanist, author of ''Flora Badensis ...
in 1788. It is found in
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
to central
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
and southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. *''F. r. albogularis'', described as ''Figulus albogularis'' by von Spix in his ''Avium species novae'' in 1824. It is found in southeast Brazil, from the states of
Goiás Goiás () is a Brazilian state located in the Center-West region. Goiás borders the Federal District and the states of (from north clockwise) Tocantins, Bahia, Minas Gerais, Mato Grosso do Sul and Mato Grosso. The state capital is Goiânia. ...
and
Bahia Bahia ( , , ; meaning "bay") is one of the 26 Federative units of Brazil, states of Brazil, located in the Northeast Region, Brazil, Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo (sta ...
to
São Paulo São Paulo (, ; Portuguese for 'Saint Paul') is the most populous city in Brazil, and is the capital of the state of São Paulo, the most populous and wealthiest Brazilian state, located in the country's Southeast Region. Listed by the GaWC a ...
. *''F. r. commersoni'', described as ''Furnarius commersoni'' by von Pelzeln in his ''Zur Ornithologie Brasiliens'' in 1868. It is found in western Brazil and eastern
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
. *''F. r. paraguayae'', described by Cherrie & Reichenberger in the '' American Museum Novitates'' journal in 1921. It is found in
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
and northern Argentina. *''F. r. schuhmacheri'', described by Laubmann in his ''Verhandlungen der Ornithologischen Gesellschaft in Bayern'' in 1933.«Beiträge zur Kenntnis des Formenkreises Furnarius rufus.»
Alfred Laubmann (German). Retrieved 18 December 2015.
It is found in southern Bolivia.


Description

The rufous hornero is a medium-sized ovenbird at 18 to 20 centimetres (7–8 in) and 31 to 58 grams (1.1–2.05 oz), with males being heavier. It has a slender and slightly decurved
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
suited to eating insects, which is horn-coloured with a length of 2.5 centimetres (1.0 in). Wings length is 10.2 centimetres (4 in), with males generally being larger. The tail is short at 7.1 centimetres (2.8 in). The measurements of the species may differ due to the
Bergmann's rule Bergmann's rule is an ecogeographical rule that states that within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer ...
. The sexes are not dimorphic in their plumage. It has a rufous brown back and pale buffy brown underparts. Its wings are pale brown except for the flight feathers which are blackish with cinnamon wingbars.


Notes


References

* * * * * * * * * * * * * * *Fraga, R.M. 1980. "The Breeding of Rufous Horneros (Furnarius rufus)" Condor, 82:58-68. *Remsen, V. (2003) ''Family Furnariidae (Ovenbirds)''. in del Hoyo J., Elliott A. & Christie D.A. (2003) ''Handbook of the Birds of the World. Volume 8. Broadbills to Tapaculos'' Lynx Edicions, Barcelona *Rodrigues, M. N., Roper, J.J., (2011) "An experimental test of the benefits of hatching asynchrony in the Rufous Hornero (Furnarius rufus)" Revista Brasileira de Ornitologia, 19:17-21. *Roper, J. J. (2005). Sexually distinct songs in the duet of the sexually monomorphic Rufous Hornero. "Journal of Field Ornithology, 76: 234-236."


External links


Rufous hornero videos
on the Internet Bird Collection

(for
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
)
Rufous hornero photo gallery
VIREO
Photo-High Res
birdfinders–"Brazil Photos" {{Taxonbar, from=Q852731 Furnarius Birds of South America Birds of Argentina Birds of Brazil Birds of Paraguay Birds of Uruguay Birds of Bolivia Birds of the Cerrado Birds of the Pantanal Birds of the Caatinga National symbols of Argentina Birds described in 1788 Taxa named by Johann Friedrich Gmelin