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Fur is a thick growth of
hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...
that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket that keeps the animal warm. The fur of mammals has many uses: protection, sensory purposes, waterproofing, and camouflaging, with the primary usage being thermoregulation. The types of hair include * ''definitive'', which may be
shed A shed is typically a simple, single-story roofed structure that is used for hobbies, or as a workshop in a back garden or on an allotment. Sheds vary considerably in their size and complexity of construction, from simple open-sided ones de ...
after reaching a certain length; * '' vibrissae'', which are sensory hairs and are most commonly whiskers; * ''pelage'', which consists of guard hairs, under-fur, and awn hair; * '' spines'', which are a type of stiff guard hair used for defense in, for example,
porcupine Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predation. The term covers two families of animals: the Old World porcupines of family Hystricidae, and the New World porcupines of family, Erethizont ...
s; * ''bristles'', which are long hairs usually used in visual signals, such as the mane of a lion; * ''velli'', often called "down fur", which insulates newborn mammals; and * '' wool'', which is long, soft, and often curly. Hair length is negligible in thermoregulation, as some tropical mammals, such as sloths, have the same fur length as some arctic mammals but with less insulation; and, conversely, other tropical mammals with short hair have the same insulating value as arctic mammals. The denseness of fur can increase an animal's insulation value, and arctic mammals especially have dense fur; for example, the musk ox has guard hairs measuring as well as a dense underfur, which forms an airtight coat, allowing them to survive in temperatures of . Some desert mammals, such as camels, use dense fur to prevent solar heat from reaching their skin, allowing the animal to stay cool; a camel's fur may reach in the summer, but the skin stays at . Aquatic mammals, conversely, trap air in their fur to conserve heat by keeping the skin dry. Mammalian coats are colored for a variety of reasons, the major selective pressures including camouflage, sexual selection, communication, and physiological processes such as temperature regulation. Camouflage is a powerful influence in many mammals, as it helps to conceal individuals from predators or prey.
Aposematism Aposematism is the advertising by an animal to potential predators that it is not worth attacking or eating. This unprofitability may consist of any defences which make the prey difficult to kill and eat, such as toxicity, venom, foul taste or ...
, warning off possible predators, is the most likely explanation of the black-and-white pelage of many mammals which are able to defend themselves, such as in the foul-smelling
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or ginge ...
and the powerful and aggressive honey badger. In arctic and subarctic mammals such as the arctic fox (''Vulpes lagopus''), collared lemming (''Dicrostonyx groenlandicus''), stoat (''Mustela erminea''), and snowshoe hare (''Lepus americanus''), seasonal color change between brown in summer and white in winter is driven largely by camouflage. Differences in female and male coat color may indicate nutrition and hormone levels, important in mate selection. Some arboreal mammals, notably primates and marsupials, have shades of violet, green, or blue skin on parts of their bodies, indicating some distinct advantage in their largely arboreal habitat due to convergent evolution. The green coloration of sloths, however, is the result of a
symbiotic Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
relationship with algae. Coat color is sometimes sexually dimorphic, as in many primate species. Coat color may influence the ability to retain heat, depending on how much light is reflected. Mammals with darker colored coats can absorb more heat from solar radiation and stay warmer; some smaller mammals, such as voles, have darker fur in the winter. The white, pigmentless fur of arctic mammals, such as the polar bear, may reflect more solar radiation directly onto the skin. The term ''pelage''first known use in English (French, from Middle French, from for 'hair', from Old French , from Latin )is sometimes used to refer to an animal's complete
coat A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
. The term fur is also used to refer to animal pelts which have been processed into leather with their hair still attached. The words ''fur'' or ''furry'' are also used, more casually, to refer to hair-like growths or formations, particularly when the subject being referred to exhibits a dense coat of fine, soft "hairs". If layered, rather than grown as a single
coat A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
, it may consist of short down hairs, long guard hairs, and in some cases, medium awn hairs. Mammals with reduced amounts of fur are often called "naked", as with the naked mole-rat, or "hairless", as with hairless dogs. An animal with commercially valuable fur is known within the fur industry as a furbearer. The use of fur as clothing or decoration is controversial;
animal welfare Animal welfare is the well-being of non-human animals. Formal standards of animal welfare vary between contexts, but are debated mostly by animal welfare groups, legislators, and academics. Animal welfare science uses measures such as longevity ...
advocates object to the trapping and killing of wildlife, and to the confinement and killing of animals on fur farms.


Composition

The modern mammalian fur arrangement is known to have occurred as far back as docodonts,
haramiyida Haramiyida ("thief" from Arabic الحرامية (al ḥarāmiyah), "thief, bandit") is a possibly polyphyletic order of mammaliaform cynodonts or mammals of controversial taxonomic affinites. Their teeth, which are by far the most common remains ...
ns and eutriconodonts, with specimens of ''
Castorocauda ''Castorocauda'' is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, ''C. lutrasimilis''. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the M ...
'', ''
Megaconus ''Megaconus'' is an extinct genus of allotherian mammal from the Middle Jurassic Tiaojishan Formation of Inner Mongolia, China. The type and only species, ''Megaconus mammaliaformis'' was first described in the journal ''Nature'' in 2013. ''Meg ...
'' and '' Spinolestes'' preserving compound follicles with both guard hair and underfur. Fur may consist of three layers, each with a different type of hair.


Down hair

Down hair (also known as underfur, undercoat, underhair or ground hair) is the bottomor innerlayer, composed of wavy or curly hairs with no straight portions or sharp points. Down hairs, which are also flat, tend to be the shortest and most numerous in the coat. Thermoregulation is the principal function of the down hair, which insulates a layer of dry air next to the skin.


Awn hair

The awn hair can be thought of as a hybrid, bridging the gap between the distinctly different characteristics of down and guard hairs. Awn hairs begin their growth much like guard hairs, but less than half way to their full length, awn hairs start to grow thin and wavy like down hair. The proximal part of the awn hair assists in thermoregulation (like the down hair), whereas the
distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
part can shed water (like the guard hair). The awn hair's thin basal portion does not allow the amount of piloerection that the stiffer guard hairs are capable of. Mammals with well developed down and guard hairs also usually have large numbers of awn hairs, which may even sometimes be the bulk of the visible coat.


Guard hair

Guard hair (overhair) is the top—or outer—layer of the coat. Guard hairs are longer, generally coarser, and have nearly straight shafts that protrude through the layer of softer down hair. The
distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
end of the guard hair is the visible layer of most mammal coats. This layer has the most marked pigmentation and gloss, manifesting as coat markings that are adapted for camouflage or display. Guard hair repels water and blocks sunlight, protecting the undercoat and skin in wet or aquatic habitats, and from the sun's ultraviolet radiation. Guard hairs can also reduce the severity of cuts or scratches to the skin. Many mammals, such as the domestic dog and cat, have a
pilomotor reflex Goose bumps, goosebumps or goose-pimples are the bumps on a person's skin at the base of body hairs which may involuntarily develop when a person is tickled, cold or experiencing strong emotions such as fear, euphoria or sexual arousal. The fo ...
that raises their guard hairs as part of a threat display when agitated.


Mammals with reduced fur

Hair is one of the defining characteristics of mammals; however, several species or breeds have considerably reduced amounts of fur. These are often called "naked" or "hairless".


Natural selection

Some mammals naturally have reduced amounts of fur. Some semiaquatic or aquatic mammals such as
cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...
s, pinnipeds and hippopotamuses have evolved hairlessness, presumably to reduce resistance through water. The naked mole-rat has evolved hairlessness, perhaps as an adaptation to their subterranean life-style. Two of the largest extant terrestrial mammals, the elephant and the rhinoceros, are largely hairless. The hairless bat is mostly hairless but does have short bristly hairs around its neck, on its front toes, and around the throat sac, along with fine hairs on the head and tail membrane. Most hairless animals cannot go in the sun for long periods of time, or stay in the cold for too long.
Marsupials Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a po ...
are born hairless and grow out fur later in development. Humans are the only primate species that have undergone significant hair loss. The hairlessness of humans compared to related species may be due to loss of functionality in the
pseudogene Pseudogenes are nonfunctional segments of DNA that resemble functional genes. Most arise as superfluous copies of functional genes, either directly by DNA duplication or indirectly by Reverse transcriptase, reverse transcription of an mRNA trans ...
KRTHAP1 (which helps produce keratin) Although the researchers dated the mutation to 240 000 ya, both the Altai Neandertal and Denisovan have the loss-of-function mutation, indicating it is much older. Mutations in the gene HR can lead to complete hair loss, though this is not typical in humans.


Artificial selection

At times, when a hairless domesticated animal is discovered, usually owing to a naturally occurring genetic mutation, humans may intentionally inbreed those hairless individuals and, after multiple generations, artificially create breeds that are hairless. There are several breeds of hairless cats, perhaps the most commonly known being the Sphynx cat. Similarly, there are some breeds of hairless dogs. Other examples of artificially selected hairless animals include the hairless guinea-pig, nude mouse, and the
hairless rat The fancy rat (''Rattus norvegicus domestica'') is the domesticated form of ''Rattus norvegicus'', the brown rat, and the most common species of rat kept as a pet. The name ''fancy rat'' derives from the use of the noun ''fancy'' for a hobby, ...
.


Use in clothing

Fur has long served as a source of clothing for humans, including Neanderthals. Historically, it was worn for its insulating quality, with aesthetics becoming a factor over time. Pelts were worn in or out, depending on their characteristics and desired use. Today fur and trim used in garments may be dyed bright colors or to mimic exotic animal patterns, or shorn close like velvet. The term "a fur" may connote a coat, wrap, or shawl. The manufacturing of fur clothing involves obtaining animal pelts where the hair is left on the animal's processed skin. In contrast, making leather involves removing the hair from the hide or pelt and using only the skin. Fur is also used to make felt. A common felt is made from beaver fur and is used in bowler hats, top hats, and high-end
cowboy hats The cowboy hat is a high-crowned, wide-brimmed hat best known as the defining piece of attire for the North American cowboy. Today it is worn by many people, and is particularly associated with ranch workers in the western and southern United ...
.Chamber's journal, Published by Orr and Smith, 1952, p. 200, Original from the University of Michigan. Common furbearers used include
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelv ...
,
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
,
mink Mink are dark-colored, semiaquatic, carnivorous mammals of the genera ''Neogale'' and '' Mustela'' and part of the family Mustelidae, which also includes weasels, otters, and ferrets. There are two extant species referred to as "mink": the A ...
,
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
,
beaver Beavers are large, semiaquatic rodents in the genus ''Castor'' native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers ar ...
, ermine,
otter Otters are carnivorous mammals in the subfamily Lutrinae. The 13 extant otter species are all semiaquatic, aquatic, or marine, with diets based on fish and invertebrates. Lutrinae is a branch of the Mustelidae family, which also includes wea ...
,
sable The sable (''Martes zibellina'') is a species of marten, a small omnivorous mammal primarily inhabiting the forest environments of Russia, from the Ural Mountains throughout Siberia, and northern Mongolia. Its habitat also borders eastern Kaza ...
,
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
, seal,
coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
,
chinchilla Chinchillas are either of two species (''Chinchilla chinchilla'' and ''Chinchilla lanigera'') of crepuscular rodents of the parvorder Caviomorpha. They are slightly larger and more robust than ground squirrels, and are native to the Andes mount ...
, raccoon, lemur, and
possum Possum may refer to: Animals * Phalangeriformes, or possums, any of a number of arboreal marsupial species native to Australia, New Guinea, and Sulawesi ** Common brushtail possum (''Trichosurus vulpecula''), a common possum in Australian urban a ...
. File:Buckingham-palace-guard-11279634947G5ru.jpg, The iconic bearskins of the Queen's Guard at Buckingham Palace are made from the fur of
American black bear The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
s. File:Carl Eielson.jpg, A seal fur coat worn by
Carl Ben Eielson Carl Benjamin "Ben" Eielson (July 20, 1897 – November 9, 1929) was an American aviator, bush pilot and explorer. Eielson Air Force Base in Alaska is named in his honor. In 1997 Carl Ben Eielson was inducted into the North Dakota Aviation Ha ...
(1897–1929), USAF pilot and Arctic explorer File:Pelz-Verkaufsstand in Tallinn, Estland.jpg, A fur store in Tallinn, Estonia, in 2019


See also

* Angora wool *
Animal coloration Animal coloration is the general appearance of an animal resulting from the reflection or emission of light from its surfaces. Some animals are brightly coloured, while others are hard to see. In some species, such as the peafowl, the male h ...
* Cat coat genetics * Coat (animal) *
Plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
* Rabbit hair * Tanning (leather)


References


External links

{{Authority control Mammal anatomy Non-timber forest products