Fulmar Formation
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The fulmars are tubenosed seabirds of the family Procellariidae. The family consists of two
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species and two extinct fossil species from the Miocene. Fulmars superficially resemble gulls, but are readily distinguished by their flight on stiff wings, and their tube noses. They breed on cliffs, laying one or rarely two eggs on a ledge of bare rock or on a grassy cliff. Outside the breeding season, they are pelagic, feeding on fish,
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
and
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
in the open ocean. They are long-lived for birds, living for up to 40 years. Historically, the northern fulmar lived on the Isle of St Kilda, where it was extensively hunted. The species has expanded its breeding range southwards to the coasts of England and northern France.


Taxonomy

The genus ''Fulmarus'' was introduced in 1826 by the English naturalist James Stephens. The name comes from the Old Norse ''Fúlmár'' meaning "foul-mew" or "foul-gull" because of the birds' habit of ejecting a foul-smelling oil. The type species was designated by George Gray in 1855 as the northern fulmar . As members of
Procellaridae The family (biology), family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prion (bird), prions, and the shearwaters. This family is part of the bird order (biology), orde ...
and then the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Procellariiformes, they share certain traits. First, they have nasal passages that attach to the upper bill called naricorns. The bills of Procellariiformes are unique in being split into between seven and nine horny plates. Finally, they produce a
stomach oil Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm petrel ...
made up of wax esters and triglycerides that is stored in the proventriculus. This can be sprayed out of their mouths as a defence against predators and as an energy rich food source for chicks and for the adults during their long flights. It will mat the
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of avian predators, which can lead to their death. Fulmars have a salt gland that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a strong saline solution from their nose.


Extant species

The genus contains the following two species.


Fossils

Two prehistoric species have been described from fossil bones found on the Pacific coast of California: ''Fulmarus miocaenus'' (
Temblor Formation The Temblor Formation is a geologic formation in California. It preserves fossils dating back from the Late Oligocene to the Middle Miocene of the Neogene period. It is notable for the famous Sharktooth Hill deposit (otherwise known as Ernst Q ...
) and ''Fulmarus hammeri'' from the Miocene.


Description

The two fulmars are closely related seabirds occupying the same niche in different oceans. The northern fulmar (''Fulmarus glacialis'') or just fulmar lives in the North Atlantic and North Pacific, whereas the southern fulmar, (''Fulmarus glacialoides'') is, as its name implies, a bird of the Southern Ocean. These
birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
look superficially like gulls, but are not closely related, and are in fact petrels. The northern species is grey and white with a yellow bill, in length with a wingspan.Maynard, B. J. (2003) The southern form is a paler bird with dark wing tips, long, with a wingspan.


Behavior


Breeding

Both recent species breed on cliffs, laying a single white egg. Unlike many small to medium birds in the Procellariiformes, they are neither
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
breeders, nor do they use
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s; their eggs are laid on the bare rock or in shallow depressions lined with plant material. In Britain, northern fulmars historically bred on St. Kilda (where their harvesting for oil, feathers and meat was central to the islands' economy). They spread into northern Scotland in the 19th century, and to the rest of the United Kingdom by 1930. The expansion has continued further south; the fulmar can now often be seen in the English Channel and in France along the northern and western coasts, with breeding pairs or small colonies in Nord,
Picardy Picardy (; Picard and french: Picardie, , ) is a historical territory and a former administrative region of France. Since 1 January 2016, it has been part of the new region of Hauts-de-France. It is located in the northern part of France. Hi ...
, Normandy and along the Atlantic coast in Brittany.Yeatman, L (1976)


Feeding

Fulmars are highly pelagic outside the breeding season, like most
tubenoses Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are oft ...
, feeding on fish, small
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
,
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
, crustaceans, marine worms, and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
. The range of these species increased greatly in the 20th century due to the availability of fish offal from commercial fleets, but may contract because of less food from this source and climatic change. The population increase has been especially notable in the British Isles.Bull, J. & Farrand Jr., J. (1993) Like other petrels, their walking ability is limited, but they are strong fliers, with a stiff wing action quite unlike the gulls. They look bull-necked compared to gulls, and have short stubby bills. They are long-lived, the longest recorded lifespan for ''F. glacialis'' being 40 years, 10 months and 16 days.


Relationship with humans

Fulmars have for centuries been hunted for food. The engraver Thomas Bewick wrote in 1804 that " Pennant, speaking of those irdswhich breed on, or inhabit, the Isle of St Kilda, says—'No bird is of so much use to the islanders as this: the Fulmar supplies them with oil for their lamps, down for their beds, a delicacy for their tables, a balm for their wounds, and a medicine for their distempers. A photograph by
George Washington Wilson George Washington Wilson (7 February 1823 – 9 March 1893) was a pioneering Scottish photographer. In 1849, he began a career as a portrait miniaturist, switching to portrait photography in 1852. He received a contract to photograph ...
taken about 1886 shows a "view of the men and women of St Kilda on the beach dividing up the catch of Fulmar".
James Fisher James Fisher may refer to: Politics *James Fisher (physician) (died 1822), Scottish-born physician and politician in Lower Canada *James Hurtle Fisher (1790–1875), South Australian lawyer, first mayor of Adelaide *James Fisher (Wisconsin politic ...
, author of ''The Fulmar'' (1952) calculated that every person on St Kilda consumed over 100 fulmars each year; the meat was their staple food, and they caught around 12,000 birds annually. Fulmar eggs were collected until the late 1920s in the St Kilda islands by their men scaling the cliffs. The eggs were buried in St Kilda peat ash to be eaten through the cold, northern winters. The eggs were considered to taste like duck eggs in taste and nourishment. However, when the human population left St Kilda in 1930, the fulmar population did not suddenly increase. Both the southern fulmar and the northern fulmar are listed as of Least Concern by the International Union for Conservation of Nature (IUCN).


Gallery

File:Northern Fulmar, Finstown, Orkney.JPG, Northern fulmar on the nest in Orkney, Scotland File:Southern Fulmar closeup.jpg, Southern fulmar in Drake's Passage. File:Fulmarus glacialis 1 8.jpg, Northern fulmar, breeding on Bear Island (Norway) File:Northern Fulmar (Fulmarus glacialis) flying, side-on.jpg, Northern fulmar in flight over Faxaflói ( Iceland) File:Fulmarus glacialis 1 2.jpg, Northern fulmar, breeding on Bear Island File:Eissturmvogel02.jpg, Northern fulmar, at the Norwegian bird-island
Runde Runde is an island in the municipality of Herøy, Møre og Romsdal, Herøy in Møre og Romsdal county, Norway. The island has a population of 113 people (as of 2015), and it is connected by the Runde Bridge to the island of Remøya to the south. ...
File:Northern Fulmar from the Crossley ID Guide Britain and Ireland.jpg, Composite image of northern fulmars in different plumages


References


Sources

* * * * * See also more recent publication(s) with similar title.


Further reading

*


External links

*
Northern fulmar profile
as part of BTO BirdFacts
Northern fulmar videos photos and sounds
on the Internet Bird Collection
Southern fulmar videos photos and sounds
on the Internet Bird Collection {{Taxonbar, from=Q311224 Fulmarus Seabirds Taxa named by James Francis Stephens