Fraxinus Americana (Oleaceae)
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''Fraxinus americana'', the white ash or American ash, is a species of
ash tree ''Fraxinus'' (), commonly called ash, is a genus of flowering plants in the olive and lilac family, Oleaceae. It contains 45–65 species of usually medium to large trees, mostly deciduous, though a number of subtropical species are evergree ...
native to eastern and central
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. The species is native to
mesophytic Mesophytes are terrestrial plants which are neither adapted to particularly dry nor particularly wet environments. An example of a mesophytic habitat would be a rural temperate meadow, which might contain Solidago, goldenrod, Trifolium, clover, Leuc ...
hardwood Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from ...
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s from
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
west to
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
, south to northern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, and southwest to eastern
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. Isolated populations have also been found in western
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
,
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
, and
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
, and the species is reportedly
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
in
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
. There are an estimated 8 billion ash trees in the United States – the majority being the white ash trees and the green ash trees.


Characteristics

The name white ash derives from the
glaucous ''Glaucous'' (, ) is used to describe the pale grey or bluish-green appearance of the surfaces of some plants, as well as in the names of birds, such as the glaucous gull (''Larus hyperboreus''), glaucous-winged gull (''Larus glaucescens''), g ...
undersides of the leaves. It is similar in appearance to the
green ash ''Fraxinus pennsylvanica'', the green ash or red ash, is a species of ash native to eastern and central North America, from Nova Scotia west to southeastern Alberta and eastern Colorado, south to northern Florida, and southwest to Oklahoma and e ...
, making identification difficult. The lower sides of the leaves of white ash are lighter in color than their upper sides, and the outer surface of the twigs of white ash may be flaky or peeling. Green ash leaves are similar in color on upper and lower sides, and twigs are smoother. White ash leaves turn yellow or red in autumn. Despite some overlap, the two species tend to grow in different locations as well; white ash is a forest tree that commonly occurs alongside
sugar maple ''Acer saccharum'', the sugar maple, is a species of flowering plant in the soapberry and lychee family Sapindaceae. It is native to the hardwood forests of eastern Canada and eastern United States. Sugar maple is best known for being the prima ...
while green ash is a
pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so m ...
that inhabits
riparian zones A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
and disturbed areas.Common Trees of the North Carolina Piedmont
''Fraxinus americana''
/ref>New Brunswick tree and shrub
''Fraxinus americana''
The white ash's compound leaves usually have 7 leaflets per leaf whereas the counts in other ash trees more often vary. Like other species in the section ''Melioides'', ''Fraxinus americana'' is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with male and female flowers produced on separate individuals.


Cultivation and uses

White ash is one of the most used trees for everyday purposes and, to keep up with high demand, is cultivated almost everywhere possible. The wood is white and quite dense (within 20% of 670 kg/m3),White Ash
''Niche Timbers''. Retrieved on 2009-07-24.
strong, and straight-grained. Its species produces an ideal, atypical dominant excurrent structured crown. It is a traditional timber of choice for production of
baseball bat A baseball bat is a smooth wooden or metal club used in the sport of baseball to hit the ball after it is thrown by the pitcher. By regulation it may be no more than in diameter at the thickest part and no more than in length. Although histor ...
s and tool handles. The wood is also favorable for furniture and
flooring Flooring is the general term for a permanent covering of a floor, or for the work of installing such a floor covering. Floor covering is a term to generically describe any finish material applied over a floor structure to provide a walking surface ...
. A study compared it to eight other different species, and it showed the highest antibacterial activity in the context of manufacturing chopping boards. Woodworkers use the timber mainly for interior uses due to high perishability in contact with ground soil. It is also used to make
lobster trap A lobster trap or lobster pot is a portable trap that traps lobsters or crayfish and is used in lobster fishing. In Scotland (chiefly in the north), the word creel is used to refer to a device used to catch lobsters and other crustaceans. A l ...
s. Since the 1950s, it has also become a popular choice for solid
electric guitar An electric guitar is a guitar that requires external amplification in order to be heard at typical performance volumes, unlike a standard acoustic guitar (however combinations of the two - a semi-acoustic guitar and an electric acoustic gui ...
bodies. It makes a serviceable
longbow A longbow (known as warbow in its time, in contrast to a hunting bow) is a type of tall bow that makes a fairly long draw possible. A longbow is not significantly recurved. Its limbs are relatively narrow and are circular or D-shaped in cross ...
if properly worked. The wood was used in ceiling fan blades from the 1970s through the mid-1980s, though cane was sometimes simulated with plastic then. It is no longer used for ceiling fan blades in most countries. White ash is not seen in cultivation as often as
green ash ''Fraxinus pennsylvanica'', the green ash or red ash, is a species of ash native to eastern and central North America, from Nova Scotia west to southeastern Alberta and eastern Colorado, south to northern Florida, and southwest to Oklahoma and e ...
, due to its preference for undisturbed forest sites away from urban pollution and soil compaction. It sometimes has been planted for its consistently reliable autumn colors, which typically are bright orange and red hues as opposed to other species of ash that produce a uniform yellow color. Cultivation of white ash differs across North American continent. For example, within the City of
Chicago (''City in a Garden''); I Will , image_map = , map_caption = Interactive Map of Chicago , coordinates = , coordinates_footnotes = , subdivision_type = Country , subdivision_name ...
region, 2010 statistics show most common street tree species is white ash at 6.2%. Along with third ranked green type at 4.9%, ashes combine to make up 11% percent of the city's street trees, with an overall population of 13,648,044 million standing ashes within
Cook County Cook County is the most populous county in the U.S. state of Illinois and the second-most-populous county in the United States, after Los Angeles County, California. More than 40% of all residents of Illinois live within Cook County. As of 20 ...
alone. Autumn Purple, or Junginger, a wild variety of American white ash selected for its purple leaf color, was discovered by
University of Wisconsin A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, t ...
horticulturist Karl Junginger of McKay Nursery in
Waterloo, Iowa Waterloo is a city in and the county seat of Black Hawk County, Iowa, United States. As of the 2020 United States Census the population was 67,314, making it the eighth-largest city in the state. The city is part of the Waterloo – Cedar Falls ...
. After its introduction in 1956, it quickly became the most popular and most expensive landscaping selection, surpassing the high priced
ginkgo ''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus within ...
,
London plane London is the capital and largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary down to the North Sea, and has been a major se ...
and white/burr oak. A related species, Biltmore ash, is sometimes treated as a variety of white ash. Other taxonomists argue that Biltmore ash is its own species (''Fraxinus biltmoreana''). North American native ash tree species are used by North American frogs as a critical food source, as leaves that fall from the trees are particularly suitable for
tadpoles A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found in ...
to feed upon in ponds (both temporary and permanent), large puddles, and other water sources. Species such as
red maple ''Acer rubrum'', the red maple, also known as swamp maple, water maple, or soft maple, is one of the most common and widespread deciduous trees of eastern and central North America. The U.S. Forest Service recognizes it as the most abundant nativ ...
, which are taking the place of ash due to the ash borer, are much less suitable for the frogs as a food source — resulting in poor frog survival rates and small frog sizes. The lack of
tannins Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', f ...
in the American ash variety makes them a good food source for frogs, but are not resistant to the ash borer. Varieties of ash from outside North America typically have much higher tannin levels and resist the borer. Maples and various non-native invasive trees, trees that are taking the place of American ash species in the North American ecosystem, typically have much higher leaf tannin levels. Ash species native to North America also provide important habitat and food for various other creatures that are native to North America.


Emerald ash borer

The
emerald ash borer The emerald ash borer (''Agrilus planipennis''), also known by the acronym EAB, is a green buprestid or jewel beetle native to north-eastern Asia that feeds on ash species. Females lay eggs in bark crevices on ash trees, and larvae feed undern ...
(''Agrilus planipennis''), also commonly known by the
acronym An acronym is a word or name formed from the initial components of a longer name or phrase. Acronyms are usually formed from the initial letters of words, as in ''NATO'' (''North Atlantic Treaty Organization''), but sometimes use syllables, as ...
EAB, is a green
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
native to Asia. In North America, the EAB is an invasive species, highly destructive to ash trees in its introduced range. The damage of this insect rivals that of
chestnut blight The pathogenic fungus ''Cryphonectria parasitica'' (formerly ''Endothia parasitica'') is a member of the Ascomycota (sac fungi). This necrotrophic fungus is native to East Asia and South East Asia and was introduced into Europe and North America ...
and
Dutch elm disease Dutch elm disease (DED) is caused by a member of the sac fungi (Ascomycota) affecting elm trees, and is spread by elm bark beetles. Although believed to be originally native to Asia, the disease was accidentally introduced into Americas, America ...
. To put its damage in perspective, the number of
chestnuts The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelated ...
killed by the
chestnut blight The pathogenic fungus ''Cryphonectria parasitica'' (formerly ''Endothia parasitica'') is a member of the Ascomycota (sac fungi). This necrotrophic fungus is native to East Asia and South East Asia and was introduced into Europe and North America ...
was around 3.5 billion chestnut trees while there are 3.5 billion ash trees in
Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...
alone.
Dutch elm disease Dutch elm disease (DED) is caused by a member of the sac fungi (Ascomycota) affecting elm trees, and is spread by elm bark beetles. Although believed to be originally native to Asia, the disease was accidentally introduced into Americas, America ...
killed only 200 million elm trees while EAB threatens 7.5 billion ash trees in the United States. The insect threatens the entire North American genus ''Fraxinus''. Since its accidental introduction into the United States and Canada in the 1990s, and its subsequent detection in 2002, it has spread to eleven states and adjacent parts of Canada. It has killed at least 50 million ash trees so far and threatens to kill most of the ash trees throughout North America. White ash exhibits a little more resistance to the EAB than green ash, which has nearly no resistance; however this could also possibly be due to white ash not being used in landscaping as extensively or placed in high-stress environments. An infested tree can be recognized by premature fall color and leaf
senescence Senescence () or biological aging is the gradual deterioration of functional characteristics in living organisms. The word ''senescence'' can refer to either cellular senescence or to senescence of the whole organism. Organismal senescence inv ...
observed on affected branches between August and last week of September. Before the EAB was officially identified, such dieback symptoms were thought to have been caused by a vascular disease classified as ash yellows. It was assumed damage was caused by the EAB taking advantage of weakened trees. Other recognizable signs regularly observed have been upper crown dieback,
epicormic shoot An epicormic shoot is a shoot growing from an epicormic bud, which lies underneath the bark of a trunk, stem, or branch of a plant. Epicormic buds lie dormant beneath the bark, their growth suppressed by hormones from active shoots higher up ...
s or sprouts, bark lesions,
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the ...
filled larval galleries, and deformed exit holes. When the "flatheaded" beetle has fully developed, it will bore out of the tree, in this instance, with a hole in a deformed "D" shape. In past it was assumed, once any symptoms had started to show, or once a tree had become infested, it was likely too late to begin management. Today only on-site professionals diagnosing an individual tree can responsibly make such determinations.


Ash fungal disease

''
Hymenoscyphus fraxineus ''Hymenoscyphus fraxineus'' () is an ascomycete fungus that causes ash dieback, a chronic fungal disease of ash trees in Europe characterised by leaf loss and crown dieback in infected trees. The fungus was first scientifically described in 200 ...
'' is a fungal disease that attacks ash trees. The disease causes leaf loss,
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
dieback, and lesions in the bark. This fungus is, for the most part, fatal, both directly and indirectly, by weakening the tree's immune system so that it is more susceptible to attacks from pests or pathogens. Because of this, younger trees are more affected, and fully mature ash trees are incapable of living long enough to reproduce. The disease has spread over most of Europe and has been confirmed in Britain, however, it has not yet been found in North America. The scale of the threat is not yet known, but is thought to be able to cause significant damage to the ash landscape of Europe and the UK.


References


External links

* {{Authority control
americana Americana may refer to: *Americana (music), a genre or style of American music *Americana (culture), artifacts of the culture of the United States Film, radio and television * ''Americana'' (1992 TV series), a documentary series presented by J ...
Hardwood forest plants Medicinal plants of North America Trees of the North-Central United States Plants described in 1753 Taxa named by Carl Linnaeus Trees of the Northeastern United States Trees of the South-Central United States Trees of the Southeastern United States Dioecious plants