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Sir Francis "Frank" Darwin (16 August 1848 – 19 September 1925) was a British
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
. He was the third son of the naturalist and scientist
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
.


Biography

Francis Darwin was born in
Down House Down House is the former home of the English naturalist Charles Darwin and his family. It was in this house and garden that Darwin worked on his theory of evolution by natural selection, which he had conceived in London before moving to Down ...
,
Downe Downe, formerly Down, () is a village in Greater London, England, located within the London Borough of Bromley but beyond the London urban sprawl. Downe is south west of Orpington and south east of Charing Cross. Downe lies on a hill, and ...
,
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it faces ...
in 1848. He was the third son and seventh child of Charles Darwin and his wife
Emma Wedgwood Emma Darwin (; 2 May 1808 – 2 October 1896) was an English woman who was the cousin marriage, wife and first cousin of Charles Darwin. They were married on 29 January 1839 and were the parents of ten children, seven of whom survived to adulth ...
. He was educated at
Clapham Clapham () is a suburb in south west London, England, lying mostly within the London Borough of Lambeth, but with some areas (most notably Clapham Common) extending into the neighbouring London Borough of Wandsworth. History Early history T ...
Grammar School. He then went to
Trinity College, Cambridge Trinity College is a constituent college of the University of Cambridge. Founded in 1546 by Henry VIII, King Henry VIII, Trinity is one of the largest Cambridge colleges, with the largest financial endowment of any college at either Cambridge ...
, first studying
mathematics Mathematics is an area of knowledge that includes the topics of numbers, formulas and related structures, shapes and the spaces in which they are contained, and quantities and their changes. These topics are represented in modern mathematics ...
, then changing to
natural science Natural science is one of the branches of science concerned with the description, understanding and prediction of natural phenomena, based on empirical evidence from observation and experimentation. Mechanisms such as peer review and repeatab ...
s, graduating in 1870. He then went to study medicine at St George's Medical School, London, earning an MB in 1875, but did not practise medicine. Darwin was married three times and widowed twice. First he married Amy Richenda Ruck in 1874, but she died in 1876 four days after the birth of their son
Bernard Darwin Bernard Richard Meirion Darwin CBE JP (7 September 1876 − 18 October 1961) a grandson of the British naturalist Charles Darwin, was a golf writer and high-standard amateur golfer. He was inducted into the World Golf Hall of Fame. Biography ...
, who was later to become a golf writer. In September 1883 he married Ellen Wordsworth Crofts (1856–1903) and they had a daughter Frances Crofts Darwin (1886–1960), a poet who married the poet
Francis Cornford Francis Macdonald Cornford (27 February 1874 – 3 January 1943) was an English classical scholar and translator known for work on ancient philosophy, notably Plato, Parmenides, Thucydides, and ancient Greek religion. Frances Cornford, his w ...
and became known under her married name. His third wife was
Florence Henrietta Fisher Florence Henrietta Darwin, Lady Darwin (''née'' Fisher, previously Maitland; 31 January 18645 March 1920), was an English playwright. Early life Florence Henrietta Fisher was born in Kensington, London, to Herbert William Fisher and his wife M ...
, daughter of
Herbert William Fisher Herbert William Fisher (30 July 1826 – 17 January 1903) was a British historian, best known for his book ''Considerations on the Origin of the American War'' (1865). Life He was born at Poulshot, Wiltshire, the son of Rev. William Fisher"The Pr ...
and widow of
Frederic William Maitland Frederic William Maitland (28 May 1850 – ) was an English historian and lawyer who is regarded as the modern father of English legal history. Early life and education, 1850–72 Frederic William Maitland was born at 53 Guilford Street, Lon ...
, whom he married in 1913, the year in which he was
knighted A knight is a person granted an honorary title of knighthood by a head of state (including the Pope) or representative for service to the monarch, the Christian denomination, church or the country, especially in a military capacity. Knighthood ...
. Her sister Adeline Fisher was the first wife of Darwin's double first cousin once removed
Ralph Vaughan Williams Ralph Vaughan Williams, (; 12 October 1872– 26 August 1958) was an English composer. His works include operas, ballets, chamber music, secular and religious vocal pieces and orchestral compositions including nine symphonies, written over ...
. Francis Darwin worked with his father on experiments dealing with plant movements, specifically
phototropism Phototropism is the growth of an organism in response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi. The cells on the plant that are farthest from the light contain a hor ...
. They co-authored ''
The Power of Movement in Plants ''The Power of Movement in Plants'' is a book by Charles Darwin on phototropism and other types of movement in plants. This book continues his work in producing evidence for his theory of natural selection. As it was one of his last books, fol ...
'' (1880) and Francis Darwin published a second expanded edition of '' Insectivorous Plants'' (1888) after his father's death. Their experiments showed that the
coleoptile Coleoptile is the pointed protective sheath covering the emerging shoot in monocotyledons such as grasses in which few leaf primordia and shoot apex of monocot embryo remain enclosed. The coleoptile protects the first leaf as well as the growing ...
of a young grass seedling directs its growth toward the light by comparing the responses of seedlings with covered and uncovered coleoptiles. These observations would later lead to the discovery of
auxin Auxins (plural of auxin ) are a class of plant hormones (or plant-growth regulators) with some morphogen-like characteristics. Auxins play a cardinal role in coordination of many growth and behavioral processes in plant life cycles and are essenti ...
. Darwin was nominated by his father to the
Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature colle ...
in 1875, and was elected as a Fellow of the Society on 2 December 1875. He was elected a
Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the judges of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural science, natural knowledge, incl ...
on 8 June 1882, the same year in which his father died. Darwin edited ''
The Autobiography of Charles Darwin ''The Autobiography of Charles Darwin'' is an autobiography by the English naturalist Charles Darwin. Darwin wrote the text, which he entitled ''Recollections of the Development of my Mind and Character'', for his family. He states that he star ...
'' (1887), and produced some books of letters from the
correspondence of Charles Darwin The British naturalist Charles Darwin corresponded with his extended family and with an extraordinarily wide range of people from all over the world. The letters, over 15,000 in all, provide many insights on issues ranging from the origins of ...
; ''The Life and Letters of Charles Darwin'' (1887) and ''More Letters of Charles Darwin'' (1905). He also edited
Thomas Huxley Thomas Henry Huxley (4 May 1825 – 29 June 1895) was an English biologist and anthropologist specialising in comparative anatomy. He has become known as "Darwin's Bulldog" for his advocacy of Charles Darwin's theory of evolution. The stor ...
's ''On the Reception of the Origin of Species'' (1887).
Cambridge University , mottoeng = Literal: From here, light and sacred draughts. Non literal: From this place, we gain enlightenment and precious knowledge. , established = , other_name = The Chancellor, Masters and Schola ...
awarded him an honorary doctorate (DSc) in 1909. He also received honorary doctorates from
Dublin Dublin (; , or ) is the capital and largest city of Republic of Ireland, Ireland. On a bay at the mouth of the River Liffey, it is in the Provinces of Ireland, province of Leinster, bordered on the south by the Dublin Mountains, a part of th ...
,
Liverpool Liverpool is a city and metropolitan borough in Merseyside, England. With a population of in 2019, it is the 10th largest English district by population and its metropolitan area is the fifth largest in the United Kingdom, with a popul ...
,
Sheffield Sheffield is a city status in the United Kingdom, city in South Yorkshire, England, whose name derives from the River Sheaf which runs through it. The city serves as the administrative centre of the City of Sheffield. It is Historic counties o ...
,
Brussels Brussels (french: Bruxelles or ; nl, Brussel ), officially the Brussels-Capital Region (All text and all but one graphic show the English name as Brussels-Capital Region.) (french: link=no, Région de Bruxelles-Capitale; nl, link=no, Bruss ...
,
St Andrews St Andrews ( la, S. Andrea(s); sco, Saunt Aundraes; gd, Cill Rìmhinn) is a town on the east coast of Fife in Scotland, southeast of Dundee and northeast of Edinburgh. St Andrews had a recorded population of 16,800 , making it Fife's fou ...
, Upsala, and
Prague Prague ( ; cs, Praha ; german: Prag, ; la, Praga) is the capital and largest city in the Czech Republic, and the historical capital of Bohemia. On the Vltava river, Prague is home to about 1.3 million people. The city has a temperate ...
. He was knighted in 1913. He is buried in Cambridge. His daughter,
Frances Cornford Frances Crofts Cornford (née Darwin; 30 March 1886 – 19 August 1960) was an English poet. Life She was the daughter of the botanist Francis Darwin and Newnham College fellow Ellen Wordsworth Crofts (1856-1903), and born into the Darw ...
, was later buried with him.


Family

His first wife, Amy Ruck, died in 1876, a few days after the birth of her son
Bernard Bernard (''Bernhard'') is a French and West Germanic masculine given name. It is also a surname. The name is attested from at least the 9th century. West Germanic ''Bernhard'' is composed from the two elements ''bern'' "bear" and ''hard'' "brav ...
, and was buried in Holy Trinity Church,
Corris Corris is a village in the county of Gwynedd, Wales, about north of the town of Machynlleth. The village lies on the west bank of the Afon Dulas (which here forms the boundary with Powys), around that river's confluence with the Afon Deri ...
, North Wales. According to a letter written by Charles Darwin to his close friend,
Joseph Dalton Hooker Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For twenty years he served as director of t ...
: " I never saw anyone suffer so much as poor Frank. He has gone to N. Wales to bury the body in a little church-yard amongst the mountains". He married his second wife, Ellen Wordsworth Crofts, in 1883. She was a Fellow and lecturer at
Newnham College Newnham College is a women's constituent college of the University of Cambridge. The college was founded in 1871 by a group organising Lectures for Ladies, members of which included philosopher Henry Sidgwick and suffragist campaigner Millicent ...
. She was a member of the
Ladies Dining Society The Ladies Dining Society was a private women's dining and discussion club, based at Cambridge University. It was founded in 1890 by the author Louise Creighton and the women's activist Kathleen Lyttelton. Its members, most of whom were marr ...
in Cambridge, together with 11 other members. She died in 1903, and is buried in the churchyard of St. Andrew's Church, Girton. In 1913, he married his third wife, Lady
Florence Henrietta Fisher Florence Henrietta Darwin, Lady Darwin (''née'' Fisher, previously Maitland; 31 January 18645 March 1920), was an English playwright. Early life Florence Henrietta Fisher was born in Kensington, London, to Herbert William Fisher and his wife M ...
, the widow of
Frederic William Maitland Frederic William Maitland (28 May 1850 – ) was an English historian and lawyer who is regarded as the modern father of English legal history. Early life and education, 1850–72 Frederic William Maitland was born at 53 Guilford Street, Lon ...
. She died in 1920 and is interred in the Parish of the Ascension Burial Ground, Cambridge, opposite the grave of Sir Francis Darwin and his daughter Frances Cornford.


Publications

*''Life and Letters of Charles Darwin'' (1880); (revised 2nd edition, 1887);2nd printing of revised edition, 1888
*''The Power of Movement in Plants'' (1880) *''The Practical Physiology of Plants'' (1894) *''Elements of Botany'' (1895)
''Rustic Sounds and Other Studies in Literature and Natural History''
(1917)


See also

*
Dorothea Pertz Dorothea Frances Matilda "Dora" Pertz FLS (14 March 1859 – 6 March 1939) was a British botanist. She co-authored five papers with Francis Darwin, Charles Darwin's son. She was made a Fellow of the Linnean Society, among the first women ad ...
with whom he coauthored five papers


References

*


Further reading

* Ayres, Peter. "The Aliveness of Plants: The Darwins at the Dawn of Plant Science" London: PIckering & Chatto, 2008. * Darwin, Francis Sacheverell. (1927). ''Travels in Spain and the East, 1808-1810''. Cambridge University press (reissued by
Cambridge University Press Cambridge University Press is the university press of the University of Cambridge. Granted letters patent by Henry VIII of England, King Henry VIII in 1534, it is the oldest university press A university press is an academic publishing hou ...
, 2009; )


External links

* * * * * {{DEFAULTSORT:Darwin, Francis English biologists English botanists Fellows of the Royal Society Fellows of the Linnean Society of London Alumni of Trinity College, Cambridge Knights Bachelor Charles Darwin biographers Darwin–Wedgwood family People from Downe 1848 births 1925 deaths Burials in Cambridgeshire Presidents of the British Science Association