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The Forrest's pika (''Ochotona forresti'') is a species of mammal in the pika family, Ochotonidae. It is found in
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
, China,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, and
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
. The summer dorsal
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
and ventral pelage are dark rufous or blackish brown, and the winter dorsal pelage is a grayish brown, slightly lighter in tone than the ventral pelage. It is a generalist herbivore. It was assessed by the
IUCN Red List of Endangered Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolog ...
as insufficiently known in 1994, as
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
in 1996, and re-assessed in 2008 as a species of 
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
.


Taxonomy

The Forrest's pika belongs to the pika family, Ochotonidae, which consists of small mammals that have short ears, forelimbs very slightly longer than hindlimbs, and no external tail. It was first described by the British
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...
Michael Rogers Oldfield Thomas Michael Rogers Oldfield Thomas (21 February 1858 – 16 June 1929) was a British zoologist. Career Thomas worked at the Natural History Museum on mammals, describing about 2,000 new species and subspecies for the first time. He was appoin ...
in 1923, who gave it the scientific name ''Ochotona forresti''. The taxonomy of the species is unclear. It was earlier assigned as a subspecies of the Moupin pika ('' Ochotona thibetana)'' by Feag and Kao in 1974, Weston in 1982, and Honacki, Kinman, and Koeppl in 1982. It was also assigned to the Royle's pika ('' Ochotona roylei)'' by Corbet in 1978, and the steppe pika (''
Ochotona pusilla The steppe pika (''Ochotona pusilla'') is a small mammal of the pika family, Ochotonidae. It is found in the steppes of southern Russia and northern Kazakhstan. General description Ochotonidae includes only one genus - ''Ochotona'', formed by ...
'') by Ellerman and Morrison-Scott in 1951. It is sympatric with the Gaoligong pika (''
Ochotona gaoligongensis The Gaoligong pika (''Ochotona gaoligongensis'') is a species of mammal in the family Ochotonidae. It is endemic to China. Many of the general physical characteristics of the pika species, are shared by Gaoligong pikas. However, the Gaoligong pik ...
''), ''Ochotona macrotis chinensis'', ''Ochotona gloveri calloceps'', Royle's pika and possibly with the Moupin pika, and allopatric with the Royle's pika. However, according to
morphometric Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
and genetic studies, the Forrest's pika is a separate species that is confined to the Gaolingong Mountains. The Gaoligong pika and black pika (''
Ochotona nigritia The black pika or silver pika (''Ochotona nigritia'') is a species of mammal in the family Ochotonidae. It was thought to be common to the Yunnan Province of China where it was first discovered in 2000, it is only known from four specimens. The ...
'') have been treated as separate species, but are now considered to be conspecific, representing a melanistic (dark) colour phase of Forrest's pika. ''O. f. duoxianglaensis'' is evidently synonymous with the Moupin pika. It may also be closely related to the Chinese red pika ('' Ochotona erythrotis'').


Description

The Forrest's pika is similar to a small hare, measuring in length, and weighing . The greatest length of the skull is , and the skull is more curved than the Moupin pika. It also has a broad
interorbital region The interorbital region of the skull is located between the eyes, anterior to the braincase. The form of the interorbital region may exhibit significant variation between taxonomic groups. In oryzomyine rodents, for example, the width, form, an ...
(the region of the skull located between the eyes, anterior to the braincase), and rectangular, broad nasals. It does not have a tail. The summer dorsal
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
and ventral pelage are dark rufous or blackish brown. The ears are rounded, measure in length, and feature dark gray spots in the back. In a few individuals, the spots form a dorsal collar on the
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...
and extend onto the face, but the forehead remains brown. The dorsal side of the ears are light chestnut in color, and have a white rim. The winter dorsal pelage is grayish brown which is slightly lighter in tone than the ventral pelage. The feet are dull white in color, the foreclaws are long, and the hindfeet are long. The 
incisive foramen In the human mouth, the incisive foramen (also known as: "''anterior palatine foramen''", or "''nasopalatine foramen''") is the opening of the incisive canals on the hard palate immediately behind the incisor teeth. It gives passage to blood ves ...
(funnel-shaped opening in the bony plate of the skull, located in the roof of the mouth, immediately behind the
incisor teeth Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
where blood vessels and nerves pass) is combined with the palatal foramina, and has a wavy edge. The Moupin's pika is similar to the Forrest's pika, but it has paler ventral pelage, shorter foreclaws, a narrower skull especially across the
cheek bone In the human skull, the zygomatic bone (from grc, ζῠγόν, zugón, yoke), also called cheekbone or malar bone, is a paired irregular bone which articulates with the maxilla, the temporal bone, the sphenoid bone and the frontal bone. It is si ...
, and buffy patches behind the ears, which do not meet around the back of the neck.


Distribution and habitat

The Forrest's pika occurs in the eastern
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
and
Kangri Garpo Kangri Karpo (), also spelt Gangri Garbo (), is a mountain range in eastern Tibet, located primarily in Nyingchi Prefecture as well as a portion of Qamdo Prefecture in the Tibet Autonomous Region, China. The mountain range lies to the east of the ...
mountains in
southwestern China Southwest China () is a region in the south of the People's Republic of China. Geography Southwest China is a rugged and mountainous region, transitioning between the Tibetan Plateau to the west and the Chinese coastal hills (东南丘陵) and ...
(northwestern
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
and southeastern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa, Taman ...
), Bhutan,
northeastern India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...
(
Arunachal Pradesh Arunachal Pradesh (, ) is a state in Northeastern India. It was formed from the erstwhile North-East Frontier Agency (NEFA) region, and became a state on 20 February 1987. It borders the states of Assam and Nagaland to the south. It shares ...
and
Sikkim Sikkim (; ) is a state in Northeastern India. It borders the Tibet Autonomous Region of China in the north and northeast, Bhutan in the east, Province No. 1 of Nepal in the west and West Bengal in the south. Sikkim is also close to the Silig ...
), and northwestern Myanmar. It is observed in the high elevation regions of northern
Arunachal Pradesh Arunachal Pradesh (, ) is a state in Northeastern India. It was formed from the erstwhile North-East Frontier Agency (NEFA) region, and became a state on 20 February 1987. It borders the states of Assam and Nagaland to the south. It shares ...
, above from sea level in the
Tawang district Tawang district (Pron:/tɑ:ˈwæŋ or təˈwæŋ/) is the smallest of the 26 administrative districts of Arunachal Pradesh state in northeastern India. With a population of 49,977, it is the eighth least populous district in the country (out o ...
, and also in high elevation regions of the
West Kameng West Kameng (pronounced ) is a district of Arunachal Pradesh in northeastern India. It accounts for 8.86% of the total area of the state. The name is derived from the Kameng river, a tributary of the Brahmaputra, that flows through the district ...
and
East Kameng East Kameng district is one of districts of Arunachal Pradesh state in northeastern, India. It shares an international border with China in the north and district borders with West Kameng district to the west, Pakke-Kessang district to the sou ...
districts. In 1997, it occurrence was reported just north of the
Upper Dibang Valley district Dibang Valley (Pron:/dɪˈbæŋ/) is a district of Arunachal Pradesh named after the Dibang River or the Talon as the Mishmis call it. It is the least populated district in India and has an area of . History In June 1980, Dibang Valley distr ...
, in Medog County, Tibet. It does not occur in present-day
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
, which belonged to the former
Undivided Assam Undivided Assam refers to the then undivided greater state of Assam in India soon after the Indian Independence in 1947 until 1963. In addition to the present-day Assam, it included the current states of Arunachal Pradesh, Meghalaya, Nagaland ...
before 1972. The Forrest's pika inhabits high altitude mixed coniferous and broadleaf forests, and shrubland thickets. It is also thought to occur around rock piles. It usually occurs on south-facing mountain slopes at elevations of above sea level. It is rarely found, and is one of the six pika species
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to central China, with no true population studies.


Behavior and ecology

The Forrest's pika is a generalist herbivore. Although very little is known about its behavior and ecology, the Forrest's pika is speculated to dig burrows. Very little or nothing is known about its reproduction.


Status and conservation

In 1994, the Forrest's pika was assessed as insufficiently known on the
IUCN Red List of Endangered Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolog ...
, and in 1996 its status was changed to
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
. It was re-assessed in 2008 and 2016, and rated as a species of 
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
because although the current state of its population trend is unclear, its population is not considered to be declining near the rate required to warrant the species a threatened status. Although no natural historic or ecological investigations have been done on the Forrest's pika, there are no known threats to its population. It is, however, possibly threatened by the loss of montane woodlands. The Chinese regional Red List has described the species as near threatened.


References


Notes


Citations


Bibliography

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Further reading

* * 黄文几,陈延熹,温业新, 中国啮齿类, 上海:复旦大学出版社, 1995 (in Chinese).


External links

* * * * {{Good article Pikas Mammals of Bhutan Mammals of Myanmar Mammals of China Mammals of India Mammals described in 1923 Taxa named by Oldfield Thomas Taxonomy articles created by Polbot