''Formicoxenus nitidulus'', or the shining guest ant, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...

in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Myrmicinae Myrmicinae is a subfamily of ants, with about 140 extant genera; their distribution is cosmopolitan. The pupae lack cocoons. Some species retain a functional sting. The petioles of Myrmicinae consist of two nodes. The nests are permanent and ...

. It is found in

Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...

Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...

Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...

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Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...

, and

Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...

. The

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed it as being a

vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...

Description

It is a very small ant, about 2–3 mm long, with a brown-red body, similar in shade to the ''

Formica ''Formica'' is a genus of ants of the family Formicidae, commonly known as wood ants, mound ants, thatching ants, and field ants. ''Formica'' is the type genus of the Formicidae, and of the subfamily Formicinae. The type species of genus ''For ...

'' ants that it nests with, but the body is noticeably shiny. The ants move about quickly on the surface of the nest, dwarfed by the much larger wood ants.

Distribution and habitat

The shining guest ant has a wide distribution in Northern Europe and Asia. Its range extends from Spain and North Italy to Scandinavia and eastern Siberia. In the British Isles it is known from Scotland, but not England, Wales or Ireland. Its habitats are wherever its host ants are found, largely in coniferous forests.

Ecology

This ant is known as the shining guest ant because it cohabits with various species of

wood ant The ''Formica rufa'' group is a subgeneric group within the genus ''Formica'', first proposed by William Morton Wheeler. This group contains the mound-building species of ''Formica'' commonly termed "wood ants" or "thatch-mound ants", which build ...

s, including ''

Formica aquilonia ''Formica aquilonia'', also called the Scottish wood ant, is a species of wood ant of the genus ''Formica'' which are widely distributed in Europe and Asia, occurring from Scandinavia in the north to Bulgaria and Italy in the south, and from the ...

'' and ''

Formica lugubris ''Formica lugubris'', also known as the hairy wood ant is commonly found in wooded upland areas across northern Eurasia. Colonies construct large thatched mound nests occupied by thousands of workers, and one or more queens. Workers look similar ...

''. The relationships between the ''Formicoxenus nitidulus'' and their hosts are poorly known. They seem to rear their own brood in a separate nest connected to their host's nest. The guest ants exit to the open air via passages from their nest, and here the individual guest ant workers solicit food from wood ant workers returning to the nest, or intercept the food while it is being transferred between these workers. There may be several guest ant nests inside one wood ant nest, each with its own queen and a few dozen workers. There is no nuptial flight for this species, reproductives attracting each other on the surface by the release of

pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...

References

External links

* {{Taxonbar, from=Q308877 Myrmicinae Hymenoptera of Europe Insects described in 1846 Taxa named by William Nylander (botanist) Taxonomy articles created by Polbot