Foreign Grain Beetle
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The foreign grain beetle (''Ahasverus advena'') is a species of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
in the family
Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vo ...
. It is related to the
sawtoothed grain beetle ''Oryzaephilus surinamensis'', the sawtoothed grain beetle, is a beetle in the superfamily ''Cucujoidea''. It is a common, worldwide pest of grain and grain products as well as chocolate, drugs, and tobacco. The species's binomial name, meaning ...
.


Description

The foreign grain beetle is approximately 2 mm ( in) in length. It can be distinguished from other grain beetles from its slight projections or knobs on each front corner of the pronotum, and its club-shaped antennae. However, other '' Ahasverus'' species look very similar and particular confusion can occur with '' Ahasverus rectus'', which is also found in grain bins in North America. The larvae are worm-like, cream-colored and often reach a length of 3 mm before pupating into darker adults. Males and females are identical in appearance both as larvae and adults. The adult is usually reddish brown, or sometimes black.


Natural history


Distribution

The foreign grain beetle is found in tropical and temperate regions. It can complete development at temperatures between 20 and 35 °C.


Behavior

At 30 °C this species can only survive if
relative humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depe ...
is at least 70%, or at least 65% at 20 °C (because the
equilibrium moisture content The equilibrium moisture content (EMC) of a hygroscopic material surrounded at least partially by air is the moisture content at which the material is neither gaining nor losing moisture. The value of the EMC depends on the material and the rela ...
of the substrate will be similar), a condition necessary for the development of
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
on which it feeds. It feeds chiefly on fungi, including
yeast Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom. The first yeast originated hundreds of millions of years ago, and at least 1,500 species are currently recognized. They are estimated to constitut ...
s. It can often be found in grain storage facilities, where it feeds on the mold growing on the grain. It is also found on other stored foodstuff, such as peanuts and dried fruits. It can be found in various moist locations where fungus develop, such as the walls of houses around plumbing systems.


Life cycle

The adult female can begin laying eggs around 3 to 4 days after emerging from the
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
. It can lay up to 8 or 12 eggs per day, but generally produces 1 to 4. Eggs are laid singly or in clusters of two or three, and they hatch in 4 to 5 days. The larval stage is completed in 11 to 19 days, and pupation takes 3 to 5 days. Larval development takes longer in drier conditions. Mated males have an average life span of 159 days, and mated females live about 208 days. Unmated beetles live longer, males up to 275 days, and females up to 300.


Impact on humans

The beetle is harmless. It may inhabit dwellings and infest products such as grains, cereals, oilseeds, dried fruit, and spices, particularly if the product is moldy. The presence of the beetle is a good indicator of damp storage conditions and spoiled food.Weaver, D. 2004. Foreign Grain Beetle - Stored Grain. High Plains IPM Guide. University of Wyoming, University of Nebraska, Colorado State University, and Montana State University. It does not damage the food itself, or any structures.


Control

The beetle is controlled in an environment by eliminating damp habitat. However, this species can develop on dry fungi so drying grain or other stored products in which it already occurs will not immediately control the beetle. Fumigants or contact insecticides may be used, but unless the moist environment is eliminated the beetle is likely to re-occur. Since the foreign grain beetle does not feed on grain or other foodstuff (except mushroom) it is technically not a pest of stored products and its presence in grain does not impede its commercialization. In the US, the legislation regulating the presence of insects in grain is the Official United States Standards for Grain, Subpart A—General Provisions, which stipulates that grain is considered infested if it contains "live weevils" or "other live insects injurious to stored grain"; and in Canada the corresponding legislation is the Canadian Grain Act, which stipulates that "receipt and marketing of infested grain (i.e., grain containing any injurious, noxious or troublesome insect or animal pest) is prohibited". Since the foreign grain beetle is not injurious to grain, its presence in grain is of little consequence, except that it is a telltale that the grain is going out of condition.


Biological control

Various insect predators and
parasitic wasps Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causin ...
attack this beetle, but since it is not a pest species it has not been the subject of a biological control program.


References


External links


Foreign Grain Beetles.
University of Minnesota, Yard & Garden Brief
Foreign Grain Beetle.
University of Kentucky College of Agriculture
Foreign Grain Beetle.
PestWeb, Government of Australia

Canadian Grain Commission
Foreign Grain Beetle.
Iowa State University, Department of Entomology * {{Taxonbar, from=Q1859507 Silvanidae Beetles described in 1832 Taxa named by Joseph Waltl