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The wildlife of Liberia consists of the
flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...
and
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
of the
Republic of Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coastâ ...
. This
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
n nation has a long Atlantic coastline and a range of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
types, with a corresponding diversity of plants and animals. Liberia is considered a biodiversity hotspot and has more intact forests characteristic of the Upper Guinea Massif than do neighbouring countries. There are 2000 species of vascular plants (including 225 tree species), approximately 140 species of mammals, and over 600 species of birds.


Geography

The country lies close to the
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can als ...
with a long coastline on the Atlantic Ocean. The terrain consists of flat or undulating coastal plains, which rise to a rolling plateau and low mountains in the northeast. A number of short rivers flow from northeast to southwest. The coastal area is characterised by swamps and
mangrove forest Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fr ...
s, while inland the tropical forests give way to
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
on the drier plateau area. The climate is
equatorial Equatorial may refer to something related to: *Earth's equator **the tropics, the Earth's equatorial region **tropical climate *the Celestial equator ** equatorial orbit **equatorial coordinate system ** equatorial mount, of telescopes * equatorial ...
with some rain all year round but the main rainy season falling between May and October. Most of the coastal area receives about of rain per year but this decreases inland, with the northern part of the country being the driest, with just half the coastal rainfall. The coastal area west of Monrovia is forested and has coastal swamps and lagoons, and there are swamps around the lagoon of Bassa Bwa, but there are no substantial wetlands inland.


Biodiversity

Liberia can be considered a
biodiversity hotspot A biodiversity hotspot is a biogeographic region with significant levels of biodiversity that is threatened by human habitation. Norman Myers wrote about the concept in two articles in ''The Environmentalist'' in 1988 and 1990, after which the co ...
and has the highest remaining portion of the forests of the Upper Guinea Massif, containing many species endemic to the region. This humid coastal forest ecosystem extends from Liberia through Cote d‘Ivoire, Ghana and Sierra Leone to Togo, and is under threat from deforestation and coastal development. Liberia has more than 2000 species of vascular plant including 225 timber tree species, 140 species of mammal, over 600 species of bird and 75 species of reptile and amphibian.


Flora

Two main types of forest are found in Liberia.
Evergreen forests An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zon ...
in the south east of the country, which has very high rainfall, are characterised by dense vegetation with trees of varying heights.
Sapo National Park Sapo National Park is a national park in Sinoe County, Liberia. It is the country's largest protected area of rainforest, was the first national park established in the country, and contains the second-largest area of primary tropical rainfor ...
exemplifies this forest type, and contains many
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
species. Moist semi-deciduous forest is found in the central and northwestern parts of the country with slightly lower rainfall. This has a more even height and a more open aspect. In 2009 it was estimated that forest cover amounted to about 32% of the country's area. There are eleven partially protected national forests. Patches of mangrove occur along the coast and near the mouth of many rivers, often lining the banks of the rivers' lower reaches. Mangrove ecosystems are important for biodiversity, as a habitat for crustaceans and young fish, for flood control and erosion control. However, mangroves are threatened by overharvesting for charcoal and firewood, and already there are few remaining areas of primary forest, you can find the forest north of tilted towers.


Fauna


Mammals

The
pygmy hippopotamus The pygmy hippopotamus or pygmy hippo (''Choeropsis liberiensis'') is a small hippopotamid which is native to the forests and swamps of West Africa, primarily in Liberia, with small populations in Sierra Leone, Guinea, and Ivory Coast. It has bee ...
occurs in riverine forests inland, for example those beside the Lofa, Cestos and
Cavalla River The Cavalla River (also known as the Cavally, the Youbou and the Diougou) is a river in West Africa running from north of Mont Nimba in Guinea, through Côte d'Ivoire, to Zwedru in Liberia, and back to the border with Côte d'Ivoire. It ends in ...
s. Liberia is its main habitat, with smaller populations in neighbouring countries. The African manatee also occurs along the coast and in many of the river systems. Other mammals found in the forests are leopards, monkeys, chimpanzees, antelopes, elephants and anteaters. The
Liberian mongoose The Liberian mongoose (''Liberiictis kuhni'') is a mongoose species native to Liberia and Ivory Coast. It is the monotypic, only member of the genus ''Liberiictis''. Phylogenetic analysis shows it is closely related to other small, social mongoos ...
is probably the rarest mammal, none having been seen for years until one was captured in 1989; it is hunted for
bushmeat Bushmeat is meat from wildlife species that are hunted for human consumption, most often referring to the meat of game in Africa. Bushmeat represents a primary source of animal protein and a cash-earning commodity for inhabitants of humid tropi ...
and classified as vulnerable by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. The
red river hog The red river hog (''Potamochoerus porcus'') or bushpig (a named also used for the '' Potamochoerus larvatus''), is a wild member of the pig family living in Africa, with most of its distribution in the Guinean and Congolian forests. It is rarel ...
is much more common. Sapo National Park hosts around 125 species of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
. These include the
African golden cat The African golden cat (''Caracal aurata'') is a wild cat endemic to the rainforests of West and Central Africa. It is threatened due to deforestation and bushmeat hunting and listed as Vulnerable on the IUCN Red List. It is a close relative o ...
, the
drill A drill is a tool used for making round holes or driving fasteners. It is fitted with a bit, either a drill or driverchuck. Hand-operated types are dramatically decreasing in popularity and cordless battery-powered ones proliferating due to ...
, seven species of monkey,
Gola malimbe The Gola malimbe or Ballman's malimbe (''Malimbus ballmanni'') is a species of bird in the family Ploceidae. Description The Gola malimbe is a black forest weaver, the males have an orange-yellow nape, bright golden-yellow crescent on the breast ...
and the Liberian mongoose. The park is also home to the
African civet The African civet (''Civettictis civetta'') is a large viverrid native to sub-Saharan Africa, where it is considered common and widely distributed in woodlands and secondary forests. It is listed as Least Concern on the IUCN Red List since 2008 ...
,
giant forest hog The giant forest hog (''Hylochoerus meinertzhageni''), the only member of its genus (''Hylochoerus''), is native to wooded habitats in Africa and is generally considered the largest wild member of the pig family, Suidae; however, a few subspecie ...
,
speckle-throated otter The spotted-necked otter (''Hydrictis maculicollis''), or speckle-throated otter, is an otter native to sub-Saharan Africa. Description The spotted-necked otter is a relatively small species, with males measuring from nose to rump, and weighin ...
, seven species of
duiker A duiker is a small to medium-sized brown antelope native to sub-Saharan Africa, found in heavily wooded areas. The 22 extant species, including three sometimes considered to be subspecies of the other species, form the subfamily Cephalophina ...
, the
water chevrotain The water chevrotain (''Hyemoschus aquaticus''), also known as the fanged deer, is a small ruminant found in tropical Africa. This is the only species in the genus ''Hyemoschus''. It is the largest of the 10 species of chevrotains, basal even-t ...
and three species of
pangolin Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (, from Ancient Greek ϕολιδωτός – "clad in scales"). The one extant family, the Manidae, has three genera: '' Manis'', '' Phataginus'', and '' Smut ...
s.


Birds

White-backed vulture As of 2016, the number of bird species recorded in Liberia is 695, some resident and some migratory, with 21 of them being globally threatened. These rare species include the
white-backed vulture The white-backed vulture (''Gyps africanus'') is an Old World vulture in the family Accipitridae, which also includes eagles, kites, buzzards and hawks. It is the most common vulture species in the continent of Africa. Description Preening at t ...
, the
hooded vulture The hooded vulture (''Necrosyrtes monachus'') is an Old World vulture in the order Accipitriformes, which also includes eagles, kites, buzzards and hawks. It is the only member of the genus ''Necrosyrtes,'' which is sister to the larger ''Gyps ...
and the
Balearic shearwater The Balearic shearwater (''Puffinus mauretanicus'') is a medium-sized shearwater in the seabird family ''Procellariidae''. ''Puffinus'' is a New Latin loanword based on the English "puffin" and its variants, that referred to the cured carcass of ...
.
Lake Piso Lake Piso, also known as Lake PisuWalters, ''Guanya Pau'', p. 79. and Fisherman's Lake,Cole, ''The Liberian Yearbook'', p. 192.Nelson, ''Liberia, a Country Study'', p. 47. is an oblongStreissguth, ''Liberia in Pictures'', p. 10. tidal lagoonFox, ...
, a coastal lagoon, is one of five wetland sites in the country which have been designated as
Ramsar sites A Ramsar site is a wetland site designated to be of international importance under the Ramsar Convention,8 ha (O) *** Permanent 8 ha (P) *** Seasonal Intermittent < 8 ha(Ts) ** Marshes on inorganic soils: *** Permanent (herb dominated) (Tp) *** P ...
. Some of the larger birds found in the rainforest are
parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoid ...
s,
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
s and
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
s.
Flamingo Flamingos or flamingoes are a type of Wader, wading bird in the Family (biology), family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas ...
es are found in the coastal swamps and lagoons. Other resident species are the
black-collared lovebird The black-collared lovebird (''Agapornis swindernianus'') also known as Swindern's lovebird is a small, 13.5 cm (5 in) long, African parrot in the genus ''Agapornis''. It is a mostly green parrot with black band on the back of its neck ...
, the
blue-headed wood-dove The blue-headed wood dove (''Turtur brehmeri''), also known as the blue-headed dove, is a medium-sized (up to long), rufous brown wood-dove with bluish-grey head, iridescent green patches on wings, reddish bill, dark brown iris, and dark red fe ...
, the
white-breasted guineafowl The white-breasted guineafowl (''Agelastes meleagrides'') is a medium-sized, up to 45 cm long, terrestrial bird of the guineafowl family. Description It has a black plumage with a small, bare, red head, white breast, long, black tail, gre ...
and the
white-necked rockfowl The white-necked rockfowl (''Picathartes gymnocephalus'') is a medium-sized bird in the family Picathartidae, with a long neck and tail. Also known as the white-necked picathartes, this passerine is mainly found in rocky forested areas at higher ...
. The pepper bird (''
Pycnonotus barbatus The common bulbul (''Pycnonotus barbatus'') is a member of the bulbul family of passerine birds. It is found in north-eastern, northern, western and central Africa. Taxonomy and systematics The common bulbul was originally described in the genus ...
'') is the
national bird This is a list of national birds, including official birds of overseas territories and other states described as nations. Most species in the list are officially designated. Some species hold only an "unofficial" status. National birds See al ...
. Around 590 species of bird have been recorded in Sapo National Park. These include the
white-breasted guineafowl The white-breasted guineafowl (''Agelastes meleagrides'') is a medium-sized, up to 45 cm long, terrestrial bird of the guineafowl family. Description It has a black plumage with a small, bare, red head, white breast, long, black tail, gre ...
and
white-necked rockfowl The white-necked rockfowl (''Picathartes gymnocephalus'') is a medium-sized bird in the family Picathartidae, with a long neck and tail. Also known as the white-necked picathartes, this passerine is mainly found in rocky forested areas at higher ...
, the
African fish eagle The African fish eagle (''Haliaeetus vocifer'') or the African sea eagle, is a large species of eagle found throughout sub-Saharan Africa wherever large bodies of open water with an abundant food supply occur. It is the national bird of Malawi, Na ...
, the
African grey parrot The grey parrot (''Psittacus erithacus''), also known as the Congo grey parrot, Congo African grey parrot or African grey parrot, is an Old World parrot in the family Psittacidae. The Timneh parrot ''(Psittacus timneh)'' once was identified a ...
, the
great blue turaco The great blue turaco (''Corythaeola cristata'') is a bird species of the family Musophagidae. At in length, it is the largest species of turaco. It has predominantly grey-blue plumage with an upright blue-black crest around high. The male and ...
, the
woodland kingfisher The woodland kingfisher (''Halcyon senegalensis'') is a tree kingfisher that is widely distributed in Africa south of the Sahara. Taxonomy The Southern Africa naturalist Carl Linnaeus included the woodland kingfisher with the binomial na ...
,
egret Egrets ( ) are herons, generally long-legged wading birds, that have white or buff plumage, developing fine plumes (usually milky white) during the breeding season. Egrets are not a biologically distinct group from herons and have the same build ...
s,
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
s,
bee-eater The bee-eaters are a group of non-passerine birds in the family Meropidae, containing three genera and thirty species. Most species are found in Africa and Asia, with a few in southern Europe, Australia, and New Guinea. They are characterised by ...
s,
roller Roller may refer to: Birds *Roller, a bird of the family Coraciidae * Roller (pigeon), a domesticated breed or variety of pigeon Devices * Roller (agricultural tool), a non-powered tool for flattening ground * Road roller, a vehicle for compa ...
s and
sunbird Sunbirds and spiderhunters make up the family Nectariniidae of passerine birds. They are small, slender passerines from the Old World, usually with downward-curved bills. Many are brightly coloured, often with iridescent feathers, particularly i ...
s.


Reptiles and amphibians

'' Bitis rhinoceros'' The
Nile crocodile The Nile crocodile (''Crocodylus niloticus'') is a large crocodilian native to freshwater habitats in Africa, where it is present in 26 countries. It is widely distributed throughout sub-Saharan Africa, occurring mostly in the central, eastern ...
and the
dwarf crocodile The dwarf crocodile (''Osteolaemus tetraspis''), also known as the African dwarf crocodile, broad-snouted crocodile (a name more often used for the Asian mugger crocodile) or bony crocodile, is an African crocodile that is also the smallest extan ...
occur in the coastal mangrove swamps but are rare, while the
West African crocodile The West African crocodile, desert crocodile, or sacred crocodile (''Crocodylus suchus'') is a species of crocodile related to – and often confused with – the larger and more aggressive Nile crocodile (''C. niloticus''). Taxonomy Compared ...
occurs further inland and is relatively common. There are a wide variety of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s,
chameleon Chameleons or chamaeleons (family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 202 species described as of June 2015. The members of this family are best known for their distinct range of colors, bein ...
s,
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...
s and
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
s, including ''
Bitis rhinoceros :''Common names: West African Gaboon viper, Gabino viperMallow D, Ludwig D, Nilson G. (2003). ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. .'' ''Bitis rhinoceros'' is ...
'', a venomous viper endemic to West Africa, slender tree snakes that feed on birds and their eggs, and blind snakes in the genus ''
Typhlops ''Typhlops'' is a genus of blind snakes in the family Typhlopidae. The genus is endemic to the West Indies. Some species which were formerly placed in the genus ''Typhlops'' have been moved to the genera ''Afrotyphlops, Amerotyphlops, Anilios, ...
''. Six species of turtle are found in Liberia, three marine and three freshwater, although one of the latter, the
forest hinge-back tortoise The forest hinge-back tortoise (''Kinixys erosa''), also known commonly as the serrated hinge-back tortoise or Schweigger's tortoise, is a species of tortoise in the family Testudinidae. The species is indigenous to the tropical forests and m ...
is largely terrestrial. With the high rainfall, amphibians are plentiful in the country, and include several genera endemic to the region including ''
Silurana ''Silurana'' is a subgenus of frogs in the family Pipidae. They are closely related the clawed frog subgenus, ''Xenopus ''Xenopus'' () (Gk., ξενος, ''xenos''=strange, πους, ''pous''=foot, commonly known as the clawed frog) is a genus ...
'', '' Hymenochirus'' and ''
Pseudhymenochirus Merlin's dwarf gray frog (''Pseudhymenochirus merlini''), or Merlin's clawed frog, is a species of frog in the family Pipidae. It is monotypic within the genus ''Pseudhymenochirus''. It is found in southern Guinea-Bissau, western Guinea, and sout ...
''. Other frogs native to Liberia include the African common toad, the
western clawed frog The western clawed frog (''Xenopus tropicalis'') is a species of frog in the family Pipidae, also known as tropical clawed frog. It is the only species in the genus ''Xenopus'' to have a diploid genome. Its genome has been sequenced, making it ...
, and the
crowned bullfrog The crowned bullfrog (''Hoplobatrachus occipitalis'') is a species of frog in the family Dicroglossidae. It is found in the Sub-Saharan Africa (Algeria, Angola, Benin, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Republic of t ...
. A number of species of
tree frog A tree frog (or treefrog) is any species of frog that spends a major portion of its lifespan in trees, known as an arboreal state. Several lineages of frogs among the Neobatrachia have given rise to treefrogs, although they are not closely relat ...
live in the foliage of trees, and the worm-like
caecilian Caecilians (; ) are a group of limbless, vermiform or serpentine amphibians. They mostly live hidden in the ground and in stream substrates, making them the least familiar order of amphibians. Caecilians are mostly distributed in the tropics of ...
'' Geotrypetes seraphini'' inhabits ants' nests.


Fish

Freshwater fish found in the lakes, rivers, streams and brackish swamps of Liberia include the African brown knifefish, the Guinean killifish, the Jeanpol's killi, the Norman's lampeye, the Rancurel's lampeye and the banded lampeye. The rough-head sea catfish occurs in brackish and marine habitats and is of interest to fisheries. Other commercially-fished species are the
Madeiran ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
and
round sardinella The round sardinella (''Sardinella aurita'') is a species of ray-finned fish in the genus ''Sardinella'' found in both sides of the Atlantic Ocean and the Mediterranean Sea. ''S. aurita'' went through a large boom in catch population around 1990 ...
s, the
Spanish mackerel Scomberomorini is a tribe of ray-finned saltwater bony fishes that is commonly known as the Spanish mackerels, seerfishes or seer fish. This fish closely resembles the King Mackerel. This tribe is a subset of the mackerel family (Scombridae) – ...
, the
bonga Bonga is a town, woreda and capital of the South West Ethiopia Peoples' Region in Ethiopia. Located in the Keffa Zone upon a hill in the upper Barta valley, it has a latitude and longitude of with an elevation of 1,714 meters above sea level. ...
, and the European anchovy, and large quantities of
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
s and
lobster Lobsters are a family (biology), family (Nephropidae, Synonym (taxonomy), synonym Homaridae) of marine crustaceans. They have long bodies with muscular tails and live in crevices or burrows on the sea floor. Three of their five pairs of legs ...
s are brought ashore.


Butterflies and moths

About 530 species of butterfly are known to be from Liberia, one of which is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
.Afrotropical Butterflies
/ref>


Conservation

Like other West African nations, Liberia is experiencing destruction of its native forests and a resulting decrease in biodiversity. The main conservation threats are the replacement of forests by
rubber plantations ''Hevea brasiliensis'', the Pará rubber tree, ''sharinga'' tree, seringueira, or most commonly, rubber tree or rubber plant, is a flowering plant belonging to the spurge family Euphorbiaceae originally native to the Amazon basin, but is now pan ...
, and more recently by
oil palm ''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm ''Elaeis guineensis'' (the species name ''guineensis'' referring to its co ...
plantations, mining, lumbering, unsustainable firewood collection, charcoal production, the introduction of alien species, slash and burn cultivation practices and the uncontrolled hunting and harvesting of wild animals and plants. An action plan, running from 2017 to 2027, has been put in place which is designed to educate the populace in order to "promote the conservation and sustenance of Liberia‘s rich biological diversity, resources and culture".


References


See also

*
Flora of Liberia The wildlife of Liberia consists of the flora and fauna of the Republic of Liberia. This West African nation has a long Atlantic coastline and a range of habitat types, with a corresponding diversity of plants and animals. Liberia is considered ...
*
Environmental issues in Liberia Environmental issues in Liberia include the deforestation of tropical rainforest, the hunting of endangered species for bushmeat, the pollution of rivers and coastal waters from industrial run-off and raw sewage, and the burning and dumping of h ...
{{WildlifeofAfrica Biota of Liberia
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...