Flesh (mycology)
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In mycology, the term trama is used in two ways. In the broad sense, it is the inner, fleshy portion of a mushroom's basidiocarp, or fruit body. It is distinct from the outer layer of tissue, known as the
pileipellis The pileipellis is the uppermost layer of hyphae in the pileus of a fungal fruit body In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowe ...
or
cuticle A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structu ...
, and from the
spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...
-bearing tissue layer known as the hymenium. In essence, the trama is the tissue that is commonly referred to as the "flesh" of mushrooms and similar fungi.Largent D, Johnson D, Watling R. 1977. ''How to Identify Mushrooms to Genus III: Microscopic Features''. Arcata, CA: Mad River Press. . pp. 60–70. The second use is more specific, and refers to the "hymenophoral trama" that supports the
hymenium The hymenium is the tissue layer on the hymenophore of a fungal fruiting body where the cells develop into basidia or asci, which produce spores. In some species all of the cells of the hymenium develop into basidia or asci, while in others some ...
. It is similarly interior, connective tissue, but it is more specifically the central layer of hyphae running from the underside of the mushroom cap to the lamella or gill, upon which the hymenium rests. Various types have been classified by their structure, including trametoid, cantharelloid, boletoid, and agaricoid, with agaricoid the most common by far. In the agarcoid type, the central trama's hyphae usually run parallel to each other, with a clear boundary area called a sub-hymenium followed by the hymenium itself on the outer layer facing the environment. The word "trama" is Latin for the " weft" or "
woof Woof may refer to: * Woof (sound), a sound made by a dog usually called a "bark" * Weft in weaving, the threads that run from side to side on a loom Music * Woof (label), a record label * "Woof" (song), by Snoop Dogg, 1998 * Woofer, a loudspea ...
" yarns in the weaving of cloth. This is related to the basidiocarp trama being "filler" tissue and that analogously the woof yarn in weaving is sometimes called "fill". Furthermore, the trama tends to be soft tissue, and in weaving, the woof yarn is not tightly stretched; it therefore need not as a rule be as strong as the
warp Warp, warped or warping may refer to: Arts and entertainment Books and comics * WaRP Graphics, an alternative comics publisher * ''Warp'' (First Comics), comic book series published by First Comics based on the play ''Warp!'' * Warp (comics), a ...
yarn.


References

Mycology Fungal morphology and anatomy {{Basidiomycota-stub