Flatwoods, pineywoods, pine savannas and longleaf pine-wiregrass ecosystem are terms that refer to an
ecological community
In ecology, a community is a group or association of populations of two or more different species occupying the same geographical area at the same time, also known as a biocoenosis, biotic community, biological community, ecological community, ...
in the southeastern
coastal plain
A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Coa ...
of North America. Flatwoods are an ecosystem maintained by
wildfire
A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
or
prescribed fire
A controlled or prescribed burn, also known as hazard reduction burning, backfire, swailing, or a burn-off, is a fire set intentionally for purposes of forest management, farming, prairie restoration or greenhouse gas abatement. A control ...
and are dominated by
longleaf pine
The longleaf pine (''Pinus palustris'') is a pine species native to the Southeastern United States, found along the coastal plain from East Texas to southern Virginia, extending into northern and central Florida. In this area it is also known as ...
(''Pinus palustris''), and
slash pine
''Pinus elliottii'', commonly known as slash pine,Family, P. P. (1990). Pinus elliottii Engelm. slash pine. ''Silvics of North America: Conifers'', (654), 338. is a conifer tree native to the Southeastern United States. Slash pine is named after ...
(''Pinus elliotii'') in the tree canopy and saw palmetto (''
Serenoa repens
''Serenoa repens'', commonly known as saw palmetto, is the sole species currently classified in the genus ''Serenoa''. It is a small palm, growing to a maximum height around . It is endemic to the subtropical and tropical Southeastern United S ...
''), gallberry (''
Ilex glabra
''Ilex glabra'', also known as Appalachian tea, dye-leaves, evergreen winterberry, gallberry, and inkberry, is a species of evergreen holly native to the coastal plain of eastern North America, from coastal Nova Scotia to Florida and west to Lo ...
'') and other flammable evergreen shrubs in the understory, along with a high diversity of herb species.
[Platt, W.J. 1999. Southeastern pine savannas. In: Anderson, R.C., Fralish, J.S. & Baskin, J. (eds.) The savanna, barren, and rock outcrop communities of North America, pp. 23- 51. Cambridge University Press, Cambridge, UK.][Peet, R.K. & Allard, D.J. 1993. Longleaf pine vegetation of the southern Atlantic and eastern Gulf Coast regions: a preliminary classification. In: Hermann, S.M. (ed.) The longleaf pine ecosystem: ecology, restoration and management, pp. 45-81.] It was once one of the dominant ecosystem types of southeastern North America. Although grasses and pines are characteristic of this system, the precise composition changes from west to east, that is, from Texas to Florida. In Louisiana, savannas even differ between the east and west side of the Mississippi River. The key factor maintaining this habitat type is recurring fire. Without fire, the habitat is eventually invaded by other species of woody plants.
[
A number of rare and endangered animals are typical of this habitat including ]red-cockaded woodpecker
The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States.
Description
The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two ...
s (''Picoides borealis''), gopher tortoises (''Gopherus polyphemus
The gopher tortoise (''Gopherus polyphemus'') is a species of tortoise in the family Testudinidae. The species is native to the southeastern United States. The gopher tortoise is seen as a keystone species because it digs burrows that provide ...
''), frosted flatwoods salamander
The frosted flatwoods salamander (''Ambystoma cingulatum'') is a small (9-13.5 cm total length), elongated species of mole salamander. It has a small, indistinct head, short legs, and a long, rounded tail. Typical coloration consists of ...
s (''Ambystoma cingulatum''), and striped newts (''Notophthalmus perstriatus
The striped newt (''Notophthalmus perstriatus'') is a species of aquatic salamander native to the southeastern United States. It is a close relative of the eastern newt, with which it shares territory, and can be distinguished from the latter b ...
''). Many rare and usual herbaceous plants are found here, particularly orchids (e.g. ''Calopogon
''Calopogon'', grass pink, is a genus of terrestrial orchids (family Orchidaceae). The generic name is from Greek and means "beautiful beard", referring to the cluster of hairs adorning the labellum. The five species are native to the eastern ...
'' species, ''Pogonia ophioglossoides
''Pogonia ophioglossoides'', the snakemouth orchid or rose pogonia, is a species of orchid occurring from central Canada to the east-central and eastern United States. It is the type species of the genus '' Pogonia''. It is pollinated by bees. Th ...
''), sedges (e.g. ''Rhynchospora
''Rhynchospora'' (beak-rush or beak-sedge) is a genus of about 400 species of sedges with a cosmopolitan distribution. The genus includes both annual and perennial species, mostly with erect 3-sided stems and 3-ranked leaves. The achenes bear a ...
'' species) and carnivorous plants (e.g. ''Sarracenia
''Sarracenia'' ( or ) is a genus comprising 8 to 11 species of North American pitcher plants, commonly called trumpet pitchers. The genus belongs to the family Sarraceniaceae, which also contain the closely allied genera '' Darlingtonia'' and ...
'' species).
A second key factor is moisture. Overall, wet pine savannas have more species than pine savannas, and the distribution of each species within a savanna is intimately connected with soil moisture regimes. Temporary ponds, and seepage areas, are therefore a critical control on plant species composition. Orchids and pitcher plant
Pitcher plants are several different carnivorous plants which have modified leaves known as pitfall traps—a prey-trapping mechanism featuring a deep cavity filled with digestive liquid. The traps of what are considered to be "true" pitcher p ...
s, for example, are associated with wetter locations. But even these wetter locations burn during dry periods, allowing regeneration of species of pitcher plant and sundew
''Drosera'', which is commonly known as the sundews, is one of the largest genera of carnivorous plants, with at least 194 species. 2 volumes. These members of the family Droseraceae lure, capture, and digest insects using stalked mucilaginous ...
.
Pineywoods are characterized by low basal area
Basal area is the cross-sectional area of trees at breast height (1.3m or 4.5 ft above ground). It is a common way to describe stand density. In forest management, basal area usually refers to merchantable timber and is given on a per hectar ...
and large widely spaced mature pine. Historically, the flatwoods were dominated by longleaf pine, which can live to be 500 years old. Large scale overharvesting
Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term app ...
in conjunction with detrimental silvicultural
Silviculture is the practice of controlling the growth, composition/structure, and quality of forests to meet values and needs, specifically timber production.
The name comes from the Latin ('forest') and ('growing'). The study of forests and wo ...
practices like replacement with faster growing loblolly pine
''Pinus taeda'', commonly known as loblolly pine, is one of several pines native to the Southeastern United States, from East Texas to Florida, and north to southern New Jersey. The wood industry classifies the species as a southern yellow pine. ...
has drastically reduced the range of the longleaf pine ecosystem. Longleaf requires frequent fires, ideally every 1–3 years, which prevent invasion of the habitat by other tree species. Decades of fire exclusion in the Southeast have contributed to the decline of this community type.[ However, with the restoration of fire, and natural flooding regimes, it is possible to restore small areas of habitat.][Keddy, P.A. 2009. Thinking big: a conservation vision for the southeastern coastal plain of North America. Southeastern Naturalist 8: 213-226.] Some private organisations are engaged in such projects. With concerted effort, several large wilderness areas could still be restored east of the Mississippi River. Some of the largest remaining areas of this habitat type are found in De Soto National Forest
De Soto National Forest, named for 16th-century Spanish explorer Hernando de Soto, is of pine forests in southern Mississippi. It is one of the most important protected areas for the biological diversity of the Gulf Coast ecoregion of North Ame ...
, Eglin Air Force Base, and Apalachicola National Forest
The Apalachicola National Forest is the largest U.S. National Forest in the state of Florida. It encompasses and is the only national forest located in the Florida Panhandle. The National Forest provides water and land-based outdoors activities ...
.[
]
See also
*South Florida pine flatwoods
The south Florida pine flatwoods are a flatwoods forest community found in central and southern Florida.
These flatwoods have open canopies of slash pine (''Pinus elliottii'' var. ''densa'') above a dense understory of low shrubs and grasses in ...
References
{{Reflist
Ecoregions of the United States