Fissidens Celticus
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''Fissidens celticus'', also known by its common name Welsh pocket-moss, is a species of
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
in the family ''
Fissidentaceae Fissidentaceae is a family of haplolepideous mosses (Dicranidae) in the order Dicranales Dicranales is an order of haplolepideous mosses in the subclass Dicranidae The Dicranidae are a widespread and diverse subclass of mosses in class Br ...
''. It was discovered in 1958 in
Pembrokeshire Pembrokeshire ( ; cy, Sir Benfro ) is a Local government in Wales#Principal areas, county in the South West Wales, south-west of Wales. It is bordered by Carmarthenshire to the east, Ceredigion to the northeast, and the rest by sea. The count ...
by A.H. Norkett and was first described as a new species by
Jean Paton Jean Annette Paton ( née Comins or Comyn; born 4 January 1929) is a British botanist, bryologist and botanical illustrator. She has written many books on the bryology of the United Kingdom and the flora of Cornwall, and described several ne ...
in 1965.


Description

''Fissidens celticus'' has shoots up to tall and wide. Each shoot has many leaves, often 10 pairs or more, of equal size. The nerve in each leaf runs all the way to the leaf tip and has a distinct bend about halfway along the leaf. Male plants and capsules are not known, and it is uncertain how the species disperses. ''Fissidens celticus'' can be distinguished from '' F. exilis'' by its longer shoots, its greater number of leaves and its lack of capsules.


Habitat

''Fissidens celticus'' grows on shaded soil banks near woodland streams, especially on bare, compact patches eroded by
floodwater A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrolog ...
. It has an altitude range of between .


Distribution

''Fissidens celticus'' is generally believed to be widespread and common within suitable localities. It is apparently
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to western Europe and there is no evidence of
population decline A population decline (also sometimes called underpopulation, depopulation, or population collapse) in humans is a reduction in a human population size. Over the long term, stretching from prehistory to the present, Earth's total human population ...
.


Within the United Kingdom

The species holotype was from Hustyn Wood,
St Breock St Breock ( kw, Nanssans) is a village and a civil parish in north Cornwall, England, United Kingdom. The spelling St Breoke was also formerly in use. Geography St Breock village is 1 mile (1.6 km) west of Wadebridge immediately to the sou ...
, Cornwall, and it is common throughout the western parts of the United Kingdom. In the United Kingdom, ''F. celticus'' often associates with ''
Calypogeia arguta ''Calypogeia'' is a genus of liverworts in the family Calypogeiaceae. It contains the following species (but this list may be incomplete): * '' Calypogeia arguta'' Mont. & Nees. * '' Calypogeia azurea'' Stotler & Crotz * '' Calypogeia fissa'' (L. ...
'', ''
Diplophyllum albicans ''Diplophyllum albicans'' is a species of liverwort The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, ...
'', '' Dicranella heteromalla'', '' Dicranella rufescens'', '' Epipterygium tozeri'', '' F. bryoides'', ''
Pellia epiphylla ''Pellia epiphylla'' (sometimes known as overleaf pellia or common pellia) is a species of thallose liverwort. It occurs in North America, Europe, North Africa and parts of Asia.Smith, A. J. E. (1989) ''The Liverworts of Britain and Ireland'', C ...
'', ''
Pohlia lutescens ''Pohlia'' is a genus of mosses in the family Mniaceae, found on all continents including Antarctica. Some of its species are native to multiple continents. The center of diversity is the Northern Hemisphere. The genus name of ''Pohlia'' is in h ...
'', and '' Pseudotaxiphyllum elegans''.


England

''Fissidens celticus'' is most commonly found in
South West England South West England, or the South West of England, is one of nine official regions of England. It consists of the counties of Bristol, Cornwall (including the Isles of Scilly), Dorset, Devon, Gloucestershire, Somerset and Wiltshire. Cities and ...
, but is also present in the
south east The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
and
north west The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
. There are records of ''F. celticus'' from
Cornwall Cornwall (; kw, Kernow ) is a historic county and ceremonial county in South West England. It is recognised as one of the Celtic nations, and is the homeland of the Cornish people. Cornwall is bordered to the north and west by the Atlantic ...
,
Devon Devon ( , historically known as Devonshire , ) is a ceremonial and non-metropolitan county in South West England. The most populous settlement in Devon is the city of Plymouth, followed by Devon's county town, the city of Exeter. Devon is ...
,
Gloucestershire Gloucestershire ( abbreviated Glos) is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn and the entire Forest of Dean. The county town is the city of Gl ...
,
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it faces ...
, and
Sussex Sussex (), from the Old English (), is a historic county in South East England that was formerly an independent medieval Anglo-Saxon kingdom. It is bounded to the west by Hampshire, north by Surrey, northeast by Kent, south by the English ...
. On the
Isle of Wight The Isle of Wight ( ) is a county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the largest and second-most populous island of England. Referred to as 'The Island' by residents, the Isle of ...
, it is present in
Parkhurst Forest Parkhurst Forest is a woodland to the north-west of Newport, Isle of Wight, England. The site is partly a site of special scientific interest. It consists of ancient woodland, relict heathland and plantation woodland. The woodland is freehold ...
, and at Briddlesford Nature Reserve. As of 2001, the only known record in
Lancashire Lancashire ( , ; abbreviated Lancs) is the name of a historic county, ceremonial county, and non-metropolitan county in North West England. The boundaries of these three areas differ significantly. The non-metropolitan county of Lancashi ...
was on the River Lune Biological Heritage Site.


Wales

The species can be found all across western
Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
, including on
Anglesey Anglesey (; cy, (Ynys) Môn ) is an island off the north-west coast of Wales. It forms a principal area known as the Isle of Anglesey, that includes Holy Island across the narrow Cymyran Strait and some islets and skerries. Anglesey island ...
.


Scotland

''Fissidens celticus'' is present in much of western
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
. The Tayvallich Juniper and Fen SSSI (near
Tayvallich Tayvallich (pronounced ; gd, Taigh a' Bhealaich ) is a small village in the Knapdale area of Argyll and Bute, in Scotland. The village name has its origins in Gaelic, and means the "house of the pass". The village is built around a sheltered harbo ...
,
Argyll and Bute Argyll and Bute ( sco, Argyll an Buit; gd, Earra-Ghàidheal agus Bòd, ) is one of 32 unitary authority council areas in Scotland and a lieutenancy area. The current lord-lieutenant for Argyll and Bute is Jane Margaret MacLeod (14 July 2020) ...
) is noted for its oceanic bryophytes, including ''F. celticus''.


Northern Ireland

In
Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...
, the only known location for ''F. celticus'' is in Ness Wood ASSI in County Londonderry.


Elsewhere

''Fissidens celticus'' has been recorded in many other countries in Europe: *Germany: in the
Leuscheid {{Infobox mountain range , name = Leuscheid , map_image = , map_caption = , pushpin_map_image = , photo = , photo_caption = , state = Rhein-Sieg-Kreis, North Rhine-Westphalia, and Altenkirchen, Rhineland-Palatinate , parent = Mittelsie ...
associating with '' F. bryoides'', '' Dicranella heteromalla'', and ''
Diplophyllum albicans ''Diplophyllum albicans'' is a species of liverwort The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, ...
''. *Ireland: one specimen the
Slieveardagh Hills The Slieveardagh Hills are a low range of hills on the County Tipperary - Kilkenny border, mainly in the Tipperary barony of Slievardagh. The highest point is Clomantagh Hill
, County Tipperary and two in
County Cork County Cork ( ga, Contae Chorcaí) is the largest and the southernmost county of Ireland, named after the city of Cork, the state's second-largest city. It is in the province of Munster and the Southern Region. Its largest market towns are ...
near
Kilworth Kilworth () is a village in north County Cork, located about 2 kilometres north of Fermoy near the river Funcheon. The M8 Cork–Dublin motorway passes nearby. Kilworth has an army camp, located on the R639 regional road between Mitchelstown a ...
; associating with '' F. bryoides'' and '' F. taxifolius''. *Spain: in the Aiako Harriak Natural Park associating with '' F. viridulus''. *
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
: in the
cantons A canton is a type of administrative division of a country. In general, cantons are relatively small in terms of area and population when compared with other administrative divisions such as counties, departments, or provinces. Internationally, t ...
of
Bern german: Berner(in)french: Bernois(e) it, bernese , neighboring_municipalities = Bremgarten bei Bern, Frauenkappelen, Ittigen, Kirchlindach, Köniz, Mühleberg, Muri bei Bern, Neuenegg, Ostermundigen, Wohlen bei Bern, Zollikofen , website ...
,
St. Gallen , neighboring_municipalities = Eggersriet, Gaiserwald, Gossau, Herisau (AR), Mörschwil, Speicher (AR), Stein (AR), Teufen (AR), Untereggen, Wittenbach , twintowns = Liberec (Czech Republic) , website ...
, and
Ticino Ticino (), sometimes Tessin (), officially the Republic and Canton of Ticino or less formally the Canton of Ticino,, informally ''Canton Ticino'' ; lmo, Canton Tesin ; german: Kanton Tessin ; french: Canton du Tessin ; rm, Chantun dal Tessin . ...
. In 2019, it was first reported in
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
(and therefore in Asia). It was found in
Bozyazı Bozyazı is a town and district of Mersin Province on the Mediterranean coast of Turkey, west of the city of Mersin. Geography Bozyazı is a remote coastal district with the Taurus Mountains as a backdrop; The roads over the mountains or along ...
, Mersin Province, associating with '' Bartramia stricta'', '' Bryum dichotomum'', '' Didymodon tophaceus'', ''
Targionia hypophylla ''Targionia'' may refer to a genus of: * Targionia (plant) ''Targionia'' is a genus of liverworts in the order Marchantiales. It the only genus in the family Targioniaceae within that order. This genus has worldwide distribution in areas ...
'' and '' Timmiella barbuloides''.


References

{{Taxonbar, from=Q17207983 Dicranales Flora of the United Kingdom Flora of Germany Flora of Ireland Flora of Spain Flora of Switzerland Plants described in 1965