''Nephrolepis cordifolia'', is a

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

native to the global tropics, including northeastern

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

. It has many common names including fishbone fern, tuberous sword fern, tuber ladder fern, erect sword fern, narrow sword fern and ladder fern, and herringbone fern. It is similar to the related fern ''

Nephrolepis exaltata ''Nephrolepis exaltata'', known as the sword fern or Boston fern, is a species of fern in the family Lomariopsidaceae (sometimes treated in the families Davalliaceae or Oleandraceae, or in its own family, Nephrolepidaceae). It is native to the A ...

''.

Description

Nephrolepis cordifolia is an evergreen fern that grows to between 40 and 80 centimeters, in extreme cases up to 1 meter. It forms an underground

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

in the form of several small

tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...

s. The

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

fronds are erect and pinnate linear to lanceolate, glandular and simple. The

rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...

bears bicolored chaff scales. The petiole is covered with bicolored pale and dark brown scales. The leaflets are entire, sessile and elongate-lanceolate. They grow up to 4.8 centimeters long and up to 0.9 cm wide. They stand at a distance of less than 1 centimeter. The sori are rounded. The spores are warty, wrinkled.

Distribution

The species is originally native to north-eastern Australia and the foothills of the

Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...

, and is considered naturalised on the central east coast of

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

. They can establish themselves terrestrially, or as an

epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...

lithophyte Lithophytes are plants that grow in or on rocks. They can be classified as either epilithic (or epipetric) or endolithic; epilithic lithophytes grow on the surfaces of rocks, while endolithic lithophytes grow in the crevices of rocks (and are also ...

, provided that the location has ideal moisture and light levels. In adequate conditions, these ferns grow readily as epiphytes, typically adhering to the bark of tree branches above a water source, where they can receive round-the-clock humidity and airflow. They can be found in a multitude of environments, from deep swamps and riverbanks to rugged outcrops and rock faces, as well as roadsides, ditches, creeks, fallen trees, and even abandoned buildings and

ruins Ruins () are the remains of a civilization's architecture. The term refers to formerly intact structures that have fallen into a state of partial or total disrepair over time due to a variety of factors, such as lack of maintenance, deliberate ...

. They love moist, shady locations; in their natural range, they are usually found in swamps (as an epiphyte) or along brooks and ditches of

coniferous forest Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ...

floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...

s (growing more terrestrially). It has been introduced into

Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...

French Polynesia )Territorial motto: ( en, "Great Tahiti of the Golden Haze") , anthem = , song_type = Regional anthem , song = " Ia Ora 'O Tahiti Nui" , image_map = French Polynesia on the globe (French Polynesia centered).svg , map_alt = Location of Frenc ...

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

, and the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

. It is also widely cultivated and distributed by humans, currently found in tropical regions of North, Central and

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

(mainly in

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

, the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...

, plus parts of

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

and

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...

), in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...

, various South Pacific islands and on the

Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...

. In the

Hawaiian Islands The Hawaiian Islands ( haw, Nā Mokupuni o Hawai‘i) are an archipelago of eight major islands, several atolls, and numerous smaller islets in the North Pacific Ocean, extending some from the island of Hawaii in the south to northernmost Kur ...

, it is known as ''kupukupu'', ''okupukupu'' or ''ni'ani'au''

Invasive species

''Nephrolepis cordifolia'' has become an

invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...

is some areas where it has been introduced. In

it is listed on the

National Pest Plant Accord The National Pest Plant Accord (NPPA) is a New Zealand agreement that identifies pest plants that are prohibited from sale and commercial propagation and distribution. The Accord initially came into effect on 1 October 2001 between regional coun ...

, which prohibits the sale, cultivation and distribution of the plant. It is listed as an invasive species in

, United States.

Gallery

File:Nephrolepis cordifolia spreading in New Zealand - 2.jpg, 'Nephrolepis cordifolia'' spreading on a forest and pasture margin in the

Waikato Waikato () is a Regions of New Zealand, local government region of the upper North Island of New Zealand. It covers the Waikato District, Waipa District, Matamata-Piako District, South Waikato District and Hamilton, New Zealand, Hamilton City ...

region in New Zealand File:Nephrolepis cordifolia tamasd02.jpg, ''Nephrolepis cordifolia'' root system showing a tuber. File:Nephrolepis cordifolia Duffii kz2.jpg, Nephrolepis cordifolia 'Duffii' at

Garfield Park Conservatory Garfield Park Conservatory, located in Garfield Park in Chicago, is one of the largest greenhouse conservatories in the United States. Often referred to as "landscape art under glass", the Garfield Park Conservatory occupies approximately insi ...

File:Nephrolepis cordifolia 01.jpg, Pot plant Nephrolepis cordifolia kz1.jpg, Nephrolepis cordifolia at the

New York Botanical Garden The New York Botanical Garden (NYBG) is a botanical garden at Bronx Park in the Bronx, New York City. Established in 1891, it is located on a site that contains a landscape with over one million living plants; the Enid A. Haupt Conservatory, ...

Nephrolepis cordifolia (L.) C.Presl (AM AK296994-3).jpg, Growing down on a planked wall Nephrolepis cordifolia (L.) C.Presl (AM AK297005-2).jpg, Growing between rocks

References

External links

''Nephrolepis cordifolia''
at the Global Invasive Species Database (

Invasive Species Specialist Group Invasive may refer to: *Invasive (medical) procedure *Invasive species *Invasive observation, especially in reference to surveillance *Invasively progressive spread of disease from one organ in the body to another, especially in reference to cancer ...

) *
''Nephrolepis cordifolia''
at Weedbusters (New Zealand)
''Nephrolepis cordifolia''
at the Center for Invasive Species and Ecosystem Health (United States) {{Taxonbar, from=Q1040814 cordifolia Flora of Nepal Flora of Australia Garden plants of Australia